Internationale Vorschriften für die Prüfung von Saatgut
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1 Règles International Internationales Rules for pour Seed les Testing Essais 2016 Semences Internationale Vorschriften für die Prüfung von Saatgut Méthos validées pour analyse sanitaire s Validated Seed Health Testing Methods semences Validierte Methon zur Gesundheitsprüfung von Saatgut 7 001a: Filterpapier-Metho Métho sur buvard pour für n la détection Nachweis 7-001a: Detection Alternaria dauci in Daucus von d Alternaria dauci dauci sur an semences Samen von Daucus carota carota (carrot) seed by blotter method (carotte) (Möhre) Inklusive Inclus Including les Änrungen changes changements and und editorial redaktionelle règles corrections et Korrekturen adopted éditoriales at verabschiet adoptées auf r lors Ordinary orntlichen la General réunion Generalversammlung Meeting annuelle 2016, générale Tallinn, 2016, 2016, Estonia Tallinn, Estonie Estland Gültig Effectives ab from 1. au Januar 1er 1 January janvier
2 Validierte Chapter Méthos 7: Methon validées Validated pour zur Seed Gesundheitsprüfung analyse Health sanitaire Testing Methods s von semences Saatgut Internationale Règles Internationales Vorschriften International pour für les die Rules Essais Prüfung for Seed von Semences Saatgut Testing Validierungsstudien Étus Validation reports validation Siehe Voir See Références. References. Literaturquellen. Les Copies copies Kopien are sont available sind disponibles erhältlich by au per secrétariat from ista.fice@ista.ch ISTA l ISTA Secretariat par courriel at durch ista.fice@ista.ch. das ISTA-Sekretariat. Kommentare, Merci Please d envoyer send comments, Empfehlungen les commentaires, suggestions or Berichte suggestions or reports zu Problemen ou problems signalement relating to this problèmes dieser method Metho sur to cette sind ISTA métho bitte Seed an das Health au responsable ISTA Committee, Seed He- du betreffend alth ISTA c/o ISTA Committee Seed Secretariat. Health über Committee, die Adresse c/o s Secrétariat ISTA-Sekretariates l ISTA. zu richten. Limitation Disclaimer responsabilité Haftungsausschluss En Whilst dépit ISTA du soin has pris taken par care l ISTA to pour ensure assurer accuracy l exactitu s Obgleich méthos methods and die et ISTA s informations mit Sorgfalt scribed auf contenues die in Richtigkeit this dans method la r scrip- in diesetion, Methonbeschreibung s ISTA méthos, shall not l ISTA liable décline angegebenen for any toute loss responsabilité or Methon damage, pour und etc. In toute resulting for mationen perte from ou dommage, achtet, use haftet this etc., sie method. résultant nicht für jeglichen l utilisation Verlust, la Schan présente métho. etc., r aus r Verwendung dieser Metho resultiert. Sicherheitsmaßnahmen Sécurité Safety precautions Der Assurez-vous Ensure sichere you are Umgang familiar connaître mit with Gefahren les hazard risques im data et mikrobiologischen and prenez take les appropriate safety und sécurité die precautions, Berücksichtigung appropriées. especially entsprechenr Il est during supposé weighing Vorsichts- que out cette mesures Labor maßnahmen, métho ingredients. est It pratiquée is insbesonre assumed dans that un während persons laboratoire r carrying Herstellung microbiologie out this test r Kulturmedien, par are s in a personnes laboratory m suitable familières Autoklavieren for s carrying principes und Auswiegen out microbiological bonnes r pratiques procedures laboratoire, sind and zu familiar gewährleisten. bonnes with pratiques principles Es wird vorausgesetzt, microbiologie, Good Labo- Bestandteile, dass et ratory s diese Practice, techniques Arbeitsschritte Good d asepsie. Microbiological in einem Tous mikrobiologischen les Practice, déchets doivent and aseptic Labor être durch traités techniques. Mitarbeiter d une Dispose manière durchgeführt appropriée all waste wern, materials (par exemple die in mit an autoclavage, n appropriate Prinzipien désinfection) way r (e.g. Guten autoclaving, et Laborpraxis, selon s disinfection) règlements Guten Mikrobiologischen and locaux in accordance sécurité. Praxis with und local r health, sterilen environmental Arbeitstechnik and vertraut safety regulations. sind. Alle Abfallstfe sind auf geeignete Weise und entsprechend r vor Ort üblichen Gesundheits-, Sicherheits- und Umweltbestimmungen zu entsorgen (z. B. durch Autoklavieren, Desinfizieren). Anmerkung Note on use zur Benutzung translations r Übersetzungen Remarque sur l utilisation s traductions Die The elektronische electronic version Version enthält International die englische, Rules französische for Seed Testing und utsche inclus Version English, r ISTA-Vorschriften. La French version and électronique German Bei versions. irgendwelchen s règles If re inclue are Fragen any les questions versions bezüglich en on r interpretation anglais, Interpretation en français r et ISTA-Vorschriftemand. English Pour ist version die toute englische question finitive Version d interprétation die version. massgebliche s règles, Version. seule la version anglaise fait en Rules, alle- foi. Herausgegeben von r: Internationalen Éditées par : Vereinigung für Saatgutprüfung (ISTA) L Association Zürichstr. 50, 8303 Internationale Bassersdorf, d Essais Schweiz Semences (ISTA) Zürichstr. 50, 8303 Bassersdorf, Suisse 2017 Published International by Seed Testing Association (ISTA) 2017 The International par l Association Seed Testing Internationale Association d Essais (ISTA) Semences (ISTA) Online Zürichstr. ISSN 50, CH-8303 Bassersdorf, Switzerland Online ISSN Titel 2017 r International englischen Originalausgabe: Seed Testing Association International (ISTA) Rules for Seed Testing Titre l édition originale anglaise : International Rules for Seed Testing Alle Online Rechte ISSN vorbehalten Kein Teil dieses Werkes darf in irgendwelcher Form or durch Tous irgendwelchem droits réservés. Verfahren, Aucune sei es partie elektronisch, cette mechanisch, publication ne durch peut Fotokopie être reproduite, or Tonaufnahme All rights dans or reserved. le système durch irgenin No part récupération anres this publication ou Verfahren être transmise may reproduziert, be reproduced, sous aucune gespeichert stored forme in or ou any par verbreitet retrieval aucun stockée moyen, wern, system or électronique, ohne transmitted vorherige mécanique, in any schriftliche form or photocopiant, by Genehmigung any means, enregistrant electronic, r Internationalen ou mechanical, autrement, Vereinigung photocopying, sans permission Saatgutprüfung. recording antérieure or orwise, par écrit without l ISTA. prior permission in writing from für ISTA a-2 Effective Gültig Effectif ab 1 1. er January janvier 2017
3 Règles Internationale Internationales Rules Vorschriften for Seed pour für les Testing die Essais Prüfung Semences von Saatgut Validierte Méthos Chapter Methon validées 7: Validated pour zur Gesundheitsprüfung analyse Seed sanitaire Health Testing s von semences Methods Saatgut 7 001a: 7-001a: : Filterpapier-Metho Detection Métho sur Alternaria buvard pour für dauci n la détection Nachweis in Daucus von carota d Alternaria (carrot) seed dauci by blotter an sur Samen semences method von Daucus Daucus carota carota (Möhre) (carotte) Wirtspflanze: Host: Daucus carota Daucus L. carota L. Krankheitserreger: Pathogen(s): Alternaria Alternaria dauci dauci (J.G.Kühn) (J.G.Kühn) J.J.Groves J.J.Groves : Daucus & Skolko, syn. carota A. syn. porri L. A. f.sp. porri dauci f.sp. (J.G.Kühn) dauci (J.G.Kühn) Neerg., & Hôte Skolko, Pathogène(s) syn. Neerg., A. syn. carotae : Alternaria A. carotae (Ellis dauci (Ellis & Langlois) (J.G.Kühn) & Langlois) Stevenson J.J.Groves Stevenson & Wellman Skolko, & Wellman syn. A. porri f.sp. dauci (J.G.Kühn) Neerg., syn. A. carotae (Ellis & Langlois) Stevenson & Wellman Erstellt durch: ISTA-PDC Method Validation Prepared by: ISTA-PDC Method Validation Préparé Sub-committee par : Sous-Comité Validation s Méthos Sub-committee l ISTA-PDC Autoren: Sheppard, J. W., Cockerell, V. Roberts, S. J., Authors: Sheppard, J.W., Cockerell, V. & Roberts, S.J. Auteurs ISTA-PDC : Sheppard, Method J.W., Validation Cockerell, Sub-committee V. Roberts, S.J. Revisionsstand Historique history la révision Version 1.0, Version 1.1, : : Definition Définition s sample la Probenumfangs taille size Version l échantillon 1.2, : Positivkontrolle Addition positive hinzugefügt; control; Version utscher common 1.2, name Name r host Wirtspflanze : Ajout add contrôle hinzugefügt positif ; ajout Version nom 1.3, commun : l hôte Neue New Abbildungen Figures 1a, 1b1a, 1b Version 1.3, 1.4, : : Berichterstattung Reporting Nouvelles results figures revised 1a, r 1b Ergebnisse überarbeitet 1.4, : «Indications s résultats» Version révisé Background Hintergrund Historique This method was originally published in ISTA Die Handbook Metho ist Seed im Health Original Testing zunächst in November veröffentlicht 1964 worn im No. métho ISTA 4 and Handbook a revised été à l origine in Seed 1987 Health éditée (Gambogi, dans Testing le 1987). ISTA im Novem- Hand It was as Cette S.3. incorporated ber book 1964 on Seed als S into Health 3 Nr. 4 Testing, Annexe und wur novembre to Chapter überprüft 1964, 7: von Seed S.3 Gambogi nhealth o 4, et révisée Testing (1987). Methods en Die 1987 Metho as (Gambogi, method wur ). in n (Sheppard La Anhang métho & zum Cockerell, a Kapitel été incorporée 2002). 7 übernommen: It has à l Annexe been Seed renumbered du Health Chapitre Testing (7-001a) 7 : Methods Méthos and slightly als d Analyse Metho modified following (Sheppard s Semences studies & Cockerell, conducted métho 2002). by Sie (Sheppard International erhielt die & Seed Coc- neue Sanitaire kerell, Health Nummer 2002). Initiative 7-001a Elle und for a été Vegetable wur renumérotée leicht Crops, veränrt (7-001a) ISF (ISHI-Veg) aufgrund et légèrement von in modifiée 1999 Studien, and die suite 2001 von à (Van s r Bilsen, étus International 2003). conduites Seed The par studies Health l International compared Initiative Seed blotter for Vegetable Health and malt Initiative Crops, agar ISF methods for Vegetable (ISHI-Veg) and conclud Crops, 1999 ISF und that (ISHI-Veg) 2001 (Van two were Bilsen, 1999 equivalent. 2003) et 2001 initiiert (Van Note Bilsen, wurn. that seeds 2003). In r can Les Studie be étus simultaneously wurn ont comparé Filterpapier- les tested for méthos und Malzagarmetho presence sur buvard Alternaria et verglichen. malt radicina agar Bei et ont using können conclu que gleichermaßen les 2 méthos genutzt étaient wern. équivalentes. Beachte: Saatgut Noter kann que parallel auch auf peuvent Alternaria être simultanément radicina mit r testées gleichen pour Metho- la pré- les same method (see method 7-002a). semences sence untersucht d Alternaria wern radicina (s. Metho avec 7-002a). la même métho (voir métho 7-002a). Treated seed Behanltes (gebeiztes) Saatgut Semences traitées Seed treatments may affect performance this test. It Behanltes should only be (gebeiztes) performed Saatgut on untreated kann die seed. Testergebnisse beeinflussen. Les traitements Es peuvent sollte nur affecter unbehanltes la performance Saatgut verwentho. wern. Elle ne vrait être appliquée que sur semences non la mé- traitées. Materials Matériel Referenzmaterial: Reference material: Referenzkulturen reference cultures or or anre or appropriatnete Materialien material geeig- Substrate: Matériel Substrat: Löschpapier blotters référence or filter : or cultures papers, Filterpapier, 90 référence mm, Durchmesser circular ou autre (e.g. Whatman 90 matériel mm (z. approprié No. B. Whatman 1 or equivalent), No. 1 or free Vergleichbares), from micro-organisms and inhibitors (3 per plate) frei Substrat von Mikroorganismen : buvards (papier und filtre), Inhibitoren p. ex. Whatman (3 Blätter n o 1 pro ou Platte). équivalent, 90 mm diamètre, exempts microorganismes 90 mm ou Petrischalen d inhibiteurs (eine (3 par pro boîte) 10 Samen). Plates: 90 mm sterile Petri dishes (one per ten seeds) Platten: Incubator: operating at 20 ±2 C, equipped with timer-controlled near-ultraviolet lights (NUV, peak at Inkubator: Boîtes Pétri Temperaturbereich : boîte Pétri stérile, 20 ±2 C, diamètre ausgestattet 90 mm mit Timer-kontrolliertem (une pour 10 semences) UV-Licht (Maximum bei 360 nm; 360 nm, e.g. colour number 08, Philips; BLB, Sylvania) Étuve z. B. : Farbnummer capable maintenir 08, Philips; une température BLB, Sylvania). 20 ±2 C, Freezer: Gefrierschrank: équipée operating proche Temperaturbereich at UV 20 contrôlés ±2 C par 20 timer ±2 C. (NUV, pic à 360 nm, p. ex. couleur n o 08, Philips ; BLB, Sylvania) Congélateur : fonctionnant à 20 ±2 C Probenvorbereitung Sample preparation Préparation l échantillon It is Es vital ist to entscheind, exclu any eine possibility Kreuzkontamination cross-contamination zwischen between verschienen seed samples. Saatgutproben This can be zu achieved verhinrn. by swabbing/ Dazu sind Il spraying est nur essentiel gereinigte equipment d exclure Gerätschaften and toute gloved possibilité hands zu verwenn with 70 contamination % und ethanol. Handschuhe The entre test zu is les tragen, carried échantillons die out mit on 70-prozentigem a semences. working sample Ceci Ethanol peut 400 être be- croisée réalisé seeds hanlt as en scribed essuyant wurn. ou in pulvérisant Section les équipements International avec l éthanol Rules Der for Test à Seed 70 wird %, Testing. mit en utilisant einem Probenumfang s gants. von 400 Samen Le durchgeführt test est réalisé wie sur es im un Kapitel échantillon r travail ISTA-Vorschriften comme beschrieben décrit ist. dans la section s Règles 400 semences, ISTA. Method Metho Méthos Critical control points are indicated by CCP. Kritische 1. Place three Kontrollpunkte 90 mm filter sind papers mit KKP in each gekennzeichnet. plate and soak Les with points sterile critiques distilled/ionized contrôle sont water. indiqués Drain par away CCP. excess In je water. Petrischale jeweils 3 Filterpapierscheiben (90 Placer Plating mm) trois legen, papiers mit sterilem filtres stilliertem/ionisiertem 90 mm diamètre dans 2.1 Wasser chacune Aseptically befeuchten. s place boîtes 10 Überschüssiges Pétri, seeds, et evenly ajoutez Wasser spaced l eau abgießen. (CCP), distillée/ on 2. Ausplattieren désionisée. surface Ôter le filter liqui paper en excès. in each plate Étalement Unter Positive sterilen control Bedingungen (reference material): 10 Körner Aseptically je Platte gleichmäßig Disposer place 2.1 seeds evenly (KKP) aseptiquement spaced auf m (CCP) Filterpapier 10 onto semences verteilen. surface bien espacées filter paper in an appropriate number plates to obtain 2.2 (CCP) Positivkontrollen dans chaque (Referenzmaterial): boîte Pétri sur la Unter surface sterilen du papier Bedingungen filtre. die Körner gleichmäßig (KKP) auf r reference culture, or plate a reference culture on 2.2 Filterpapieroberfläche Contrôle positif (matériel in einer référence) entsprechenn : Placer Anzahl les media. The number plates required will pend on von semences Petrischalen manière verteilen, aseptique um eine et régulièrement Referenzkultur disposées (CCP) or sur la eine surface Referenzkultur du papier filtre auf et m répéter Medium avec zu level contamination positive-control seed erhalten, lot. ausplattieren. un nombre approprié Die Anzahl boîtes r erforrlichen pour obtenir Petrischalen la culture 3. Incubate référence, wird von for 3 r ou d at Stärke déposer 20 ±2 r C une Kontamination in culture dark. référence r Positivkontrollpartie sur 4. milieu. Transfer Le plates nombre abhängen. to freezer boîtes and requis maintain dépend at du 20 niveau ±2 C 3. contamination 3 for Tage 24 h. bei 20 ±2 du C lot im Dunkeln semences inkubieren. servant contrôle 4. Platten positif. anschließend für 24 h bei 20 ±2 C in einem 3. Gefrierschrank Incuber pendant lagern. 3 jours à 20 ±2 C à l obscurité. 4. Transférer les boîtes au congélateur et maintenir à 20 ±2 C pendant 24 h a: : Filterpapier-Metho Alternaria Métho sur dauci buvard on Daucus pour für Alternaria carota (Carrot) dauci an sur by blotter Daucus carota method carota (Möhre) (carotte) Gültig Effective Effectif ab 1 er 1. janvier January a-3
4 Validierte Chapter Méthos 7: Methon validées Validated pour zur Seed Gesundheitsprüfung analyse Health sanitaire Testing Methods s von semences Saatgut Internationale Règles Internationales Vorschriften International pour für les die Rules Essais Prüfung for Seed von Semences Saatgut Testing 7 001a: : Filterpapier-Metho Alternaria Métho sur dauci buvard on Daucus pour für Alternaria carota (Carrot) dauci an sur by blotter Daucus carota method carota (Möhre) (carotte) 5. Nach Après After r freezing, congélation, Kältebehandlung incubate incuber for pendant Inkubation 6 d 620 jours ±2 r C à Platten 20 with ±2 für alternating avec 6 Tage une bei alternance 1220 h ±2 periods C unter périos darkness wechselnn 12 and h d obscurité Lichtbedin- NUV light et C gungen: (ISTA,1984; proche-uv 12 h (NUV)(ISTA, Tempe, dunkel und 1968). anschließend 1984 Plates ; Tempe, should 12 h 1968). be UV-Licht approx. Les boîtes (ISTA, 25 cm vraient below 1984; Tempe, être lights environ 1968). and should 25 Platten cm not en sollten be ssous stacked. ungefähr s lumières 25 Examine cm unterhalb et ne seeds vraient unr Lampen pas a stereoscopic être stehen empilées. und microscope nicht gestapelt at Observer wern. 30 for fungal les semences growth and au up microscope to 80 magnification 30 pour for la 6. Die croissance intification Körner du mit mycélium Hilfe conidia. eines et Conidiophores Stereomikroskops jusqu à grossissement are bei simple 30-facher pour slightly Vergrößerung l intification branched (Fig. auf s pilzliches 1), conidies. arising Wachstum Les singly conidiophores or in untersu- small 80 or chen sont groups simples und from bei ou 80-facher peu surface ramifiés Vergrößerung (Fig. seed 1), apparaissant or on die aerial Konidien seuls mycelium. petits Conidia groupes Die are Konidienträger à usually la surface solitary, s semences sind obclavate, einfach ou sur up or to le intifizieren. ou schwach mycélium 450 µm long verzweigt aérien. (including Les (Abb. conidies beak), 1), entspringen sont pale olivaceous général einzeln isolées, brown or in en at kleinen first, forme becoming Gruppen massue, brown auf mesurant r with Samenoberfläche jusqu à age, with 450 a μm long or long pale aus m (incluant Luftmyzel. le bec), Sporen brun olive sind pâle meist au einzeln, départ et keulenförmig, brunes bis en zu vieillissant 450 µm lang avec (inkl. un bec Fortsatz), long et zunächst pâle mesurant blass venant beak up to 3 times length body (Ellis, 1971). Groups sunken conidia are sometimes visible by jusqu à oliv-braun, 3 fois im la Alter longueur braun du wernd, corps haben la conidie einen langen (Ellis, emerging clusters ir bright long beaks (Fig. 1c). 1971). blassen Des Fortsatz, groupes r bis conidies zu dreimal cachés so lang sont ist, parfois wie r visibles eigentliche grâce Sporenkörper à l extrémité (Ellis, leur long 1971). bec Gruppen clair (Fig. einge- 1c). Compare with positive control. Record number infected seeds in each plate (CCP). sunkener Comparer Konidien avec le témoin können positif. manchmal Noter an le n nombre Büscheln langer semences Fortsätze infectées erkannt dans chaque wern boîte (Abb. (CCP). 1c). Mit Positivkontrolle vergleichen. Anzahl infizierter Samen pro General Platte erfassen methods (KKP). Méthos générales Checking tolerances: Tolerances provi a means assessing wher s tolérances Methon or not : variation Les tolérances in results fournissent within Allgemeine Vérification or s between moyens tests d évaluer are sufficiently si la variation wi du as résultat to raise dans doubts Überprüfung entre about les essais accuracy r est Toleranz: suffisante results. Toleranzen pour Suitable s assurer erlauben tolerances, l exactitu which Abschätzung, can s be résultats. applied ob die Des to Variation most tolérances direct r seed Ergebnisse appropriées, health tests, eines qui eine peuvent can Testes be or found être zwischen appliquées in Table verschienen à 5B la Part plupart 1 s Tests Chapter essais ausreichend d analyse 5 sanitaire ISTA groß ist Rules, or directs, or nicht, Table peuvent um G1 an in être r Miles trouvées Genauigkeit (1963). dans r le Tableau Ergebnisse Partie zu zweifeln. 1 du chapitre Eine 5 Toleranztabelle, s Règles ISTA, die ou für dans die la 5B Reporting Tableau meisten G1 direkten results: du Miles Saatgut-Gesundheitsprüfungen The (1963). result a seed health angewent indicate wern kann, scientific fint sich name in Tabelle 5B pathogen Teil 1 test should Indication tected s Kapitels s and résultats 5 r test ISTA-Vorschriften, method : Le résultat used. d un When or essai reported in Tabelle qualité G1 ISTA in sanitaire Miles Certificate, (1963). s semences results are vrait entered indiquer unr le Or nom on an Determinations. scientifique du pathogène détecté et la métho d essai Berichterstattung The employée. report Les must résultats r indicate Ergebnisse: sont écrits number Der dans Ergebnisbericht «seeds Autres tested. déterminations einer Gesundheitsprüfung case» dans a negative le Bulletin result von ISTA. (pathogen Saatgut sollte not tected), n wis- In Le senschaftlichen rapport results doit must indiquer Name be reported s le nombre Krankheitserregers as not tected. semences und testées. die In Dans verwente le case d un Metho a positive résultat enthalten. result, négatif (pathogène Bei report Berichterstattung should non détecté)cate auf einem les résultats percentage ISTA-Bericht doivent infected sind être die rapportés seeds. Ergebnisse comme unter «Wei- non indi- détecté tere Untersuchungsergebnisse». einzutragen. Dans Der Prüfbericht le cas d un résultat muss die positif, Anzahl le rapport untersuchter doit indiquer Samen Quality le angeben. pourcentage assurance semences infectées. Im Falle eines negativen Ergebnisses (Krankheitserreger nicht nachgewiesen), müssen die Ergebnisse als Specific training Assurance nicht nachgewiesen qualité berichtet wern. Im Falle eines positiven Ergebnisses sollte r Prüfbericht n Prozentsatz spécifique infizierter Samen angeben. This test should only be performed by persons who have Formation been trained in fungal intification or unr direct supervision Ce test ne vrait someone être effectué who has. que par s personnes formées à d intification fongique ou sous la supervision Qualitätssicherung personnes qui l ont été. Spezifische Ausbildung Der Test sollte nur von Personen durchgeführt wern, die entsprechen Erfahrungen bei r Intifizierung von Pilzen Critical Points haben critiques control or unter points r direkten contrôle (CCP) Anleitung (CCP) einer Person erfolgen, die die entsprechenn Kenntnisse besitzt. La Spreading croissance hyphae s hyphes may lead peut to amener contamination à la contamination seeds. d autres Seeds semences. must refore Les semences be spaced doivent at least donc 20 or Kritische mm être espacées from Kontrollpunkte each au or moins with 20 not mm more (KKP) les than unes 10 s seeds autres per et 90 ne pas mm placer Petri dish plus (Step 102). semences par boîte Pétri 90 Die Samples mm wachsenn (étape may be 2). difficult Hyphen to können examine zu einer due to Kontamination Les contaminants, échantillons anrer Samen peuvent especially führen. être Deshalb Alternaria difficiles muss à noter tenuis, r à Abstand and/or growth r cause A. radicina. einzelnen la présence Experience Körner auf saprophytes and m great Filterpapier care particulièrement is required wenigstens for 20 Alternaria mm tection betragen. tenuis, all Desweiteren et/ou occurrences A. radicina. nicht (ISTA mehr De 1984)(Step l expérience als 10 Samen 6). et pro Supplementary une Petrischale gran attention (90 examination mm) sont auslegen requises may (Schritt be pour required la détection 2) d Aufgrund after tout ce plating qui s peut Wachstums seeds se produire. to allow von (ISTA hampered anren 1984)(étape Pilzen or eper 6). ist bisweilen Une notation die Auswertung supplémentaire erschwert. peut être Häufig nécessaire treten 13 hier inoculum to exceed contaminants by its fructification Alternaria 14 jours après tenuis le dépôt und/or s semences A. radicina pour auf. permettre Erfahrung à (Hewett, 1964)(Step 6). und inoculum, große Sorgfalt gêné par ist daher les contaminants bei r Bestimmung ou prond, erforrlich fructifier (ISTA (Hewett, 1984)(Schritt 1964)(étape 6). 6). Eine zusätzliche Untersuchung 13 bis 14 Tage nach References m Auslegen r Samen kann erforrlich sein, um Références anhand r Fruktifikation auch noch im Wachstum verlangsamte Ellis, M. B. (1971). or im Dematiaceous Korn tiefer angesielte Hyphomycetes. Pathogene Common Ellis, nachzuweisen M. wealth B. (1971). Mycological (Hewett, Dematiaceous 1964)(Schritt Institute, Hyphomycetes. Kew, 6). Surrey, Common wealth 609 pp. Mycological Institute, Kew, Surrey, Eng- England. Gambogi, land. 609 P. (1987). pp. International Seed Testing Association, Literaturquellen Gambogi, Handbook P. (1987). on Seed International Health Testing, Seed Working Testing Association, Sheet No. 4 Handbook (2nd Ed.): Alternaria on Seed Health dauci Testing, on Daucus Working carota. Sheet International (2nd M. Ed.): Seed B. (1971). Alternaria Testing Dematiaceous Association, dauci on Daucus Zurich, Hyphomycetes. carota. Switzerland. Interna- Com- No. 4 Ellis, Hewett, mon tional wealth P. Seed D. Testing (1964). Mycological Association, Testing Institute, carrot Zurich, Kew, seed Switzerland. infected Surrey, England. Alternaria P. 609 D. pp. (1964). porri f. Testing sp. dauci. carrot Proceedings seed infected with with Hewett, Gambogi, International Alternaria P. (1987). porri Seed International f. Testing sp. dauci. Association, Seed Proceedings Testing 29, Association, International Handbook Seed on Seed Testing Health Testing Testing, Association, Working (1984). 29, Sheet Report No. 4 International (2nd18th Ed.): International Seed Alternaria Testing dauci Seminar Association on on Daucus Seed (1984). Pathology, carota. Report International 18th 1984, Seed International Testing Washington Association, Seminar State on Zurich, University, Seed Pathology, Switzerland. Puyallup, July, Hewett, U.S.A. July, P. 1984, D. (1964). Washington Testing State carrot University, seed infected Puyallup, with Miles, U.S.A. Alternaria R. (1963). porri Hand f. sp. book dauci. tolerances Proceedings and measures S. R. precision (1963). Seed Hand for Testing seed book Association, testing. tolerances Proceedings 29, and mea- Miles, International International International sures precision Seed Seed Testing for Testing seed Association testing. Association, Proceedings (1984). 28 Report (3), International 18th International Seed Testing Seminar Association, on Seed Pathology, 28 (3), Roberts, July, 686. S. 1984, J., Phelps, Washington K., Taylor, State J. University, D. & Ridout, Puyallup, M. S. Roberts, U.S.A. (1993). S. J., Design Phelps, and K., interpretation Taylor, J. D. & Ridout, seed health M. S. Miles, assays. (1993). S. R. In Design (1963). Proceedings Hand book interpretation First tolerances ISTA Plant seed and Disease health measures assays. In precision Proceedings Symposium for seed on Seed testing. First Health ISTA Proceedings Testing, Plant Ottawa, Disease Committee Canada International Committee (ed. Symposium J. Seed W. Sheppard), Testing on Seed Association, pp. Health , Testing, 28 Agriculture (3), Ottawa, 525 Canada, 686. Canada (ed. Ottawa, J. W. Canada. Sheppard), pp , Agriculture Roberts, Canada, S. Ottawa, J., Phelps, Canada. K., Taylor, J. D. & Ridout, M. S. Sheppard, J. W. & Cockerell, V. (2002). Detection Sheppard, (1993). J. Design W. and Cockerell, interpretation V. (2002). Detection seed health Alternaria dauci on Daucus carota (Carrot). International Rules for Seed Testing. Annexe to Chapter 7: assays. Alternaria In Proceedings dauci on Daucus First carota ISTA (Carrot). Plant Disease International Rules Symposium for Seed on Testing. Seed Health Annexe Testing, to Chapter Ottawa, 7: Committee Seed Health Testing Methods Seed Canada Health (ed. Testing J. W. Sheppard), Methods. pp , Agriculture Tempe, J. (1968). The quantitative effect light and Tempe, Canada, J. Ottawa, (1968). Canada. The quantitative effect light and Sheppard, moisture moisture J. on on W. carrot carrot & Cockerell, seed seed infections infections V. (2002). in in blotter blotter Detection medium. medium. Proceedings Proceedings Alternaria dauci on International International Daucus carota Seed Seed (Carrot). Testing Testing International Asso Asso ciation, ciation, 33, 33, Rules for Seed Testing. Annexe to Chapter 7: Seed Health Testing Methods Tempe, J. (1968). The quantitative effect light and moisture on carrot seed infections in blotter medium. Proceedings International Seed Testing Asso ciation, 33, a-4 Effective Gültig Effectif ab 1 1. er January janvier 2017
5 Règles Internationale Internationales Rules Vorschriften for Seed pour für les Testing die Essais Prüfung Semences von Saatgut Validierte Méthos Chapter Methon validées 7: Validated pour zur Gesundheitsprüfung analyse Seed sanitaire Health Testing s von semences Methods Saatgut a b c Figure 1. a Conidies Conidia d Alternaria dauci on sur simple conidiophores or slightly simples branched ou faiblement conidiophores. ramifiés. b Single b Conidies conidia. simples. c Conidia c Conidies A. dauci d A. on dauci simple sur or conidiophores slightly branched simples conidiophores ou faiblement borne ramifiés on a single se développant rootlet initial. sur 80. une d Conidium ébauche and radicelle. simple conidiophores. 80. d Conidies 350. et conidiophores simples Validierungsstudien Étus Validation validation reports Van Bilsen, J. G. P. M. (2003). Report a comparative test Van on Bilsen, Alternaria J. G. P. dauci M. (2003). and Alternaria Report radicina a comparative on carrot test seed. on Alternaria ISTA Method dauci Validation and Alternaria Reports radicina (submitted). on carrot seed. ISTA Method Validation Reports (submitted). d Abb. 1. a Konidien von Alternaria dauci auf einfache or schwach verzweigte Konidienträger. b Einzelne Konidien. c Konidien von A. dauci an einem einfachen or schwach verzweigten Träger auf einer Keimwurzel. 80. d Konidie und Konidienträger a: : Filterpapier-Metho Alternaria Métho sur dauci buvard on Daucus pour für Alternaria carota (Carrot) dauci an sur by blotter Daucus carota method carota (Möhre) (carotte) Gültig Effective Effectif ab 1 er 1. janvier January a-5
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