Feed Efficiency in Beef Finishing Systems

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Feed Efficiency in Beef Finishing Systems M. McGee Teagasc, Animal & Grassland Research and Innovation Centre, Grange, Dunsany, Co. Meath Teagasc-IGFA Nutrition Conference Portlaoise June 2014

Feed costs Conversion of feed to product Providing feed - particularly during the indoor winter period - is the LARGEST variable cost in BEEF production: 70%+ Variable costs Grange suckler calf-to-beef system 65% 35% feed other

Annual Feed Budgets (/cow unit) Suckler calf-to-beef: 24 mth steer 8% DMI (incl. land charge Grazed grass 17% 27% 65% Grass silage Concentrates 44% 39% Optimising the contribution of grazed grass to the lifetime intake of feed and provide silage & concentrates as efficiently and at as low as cost as feasible

Profit = Income - Costs Feed costs Cow Herd Progeny Improvement in feed efficiency is worth 4-8 times more than an equivalent increase in growth rate (Gibb and McAllister, 1999; Okine et al., 2004).

High maternal cost to Suckler beef production 100% 80% 60% 40% 20% 0% Cow Feed Requirements Progeny Cow In beef production Maintenance Reqs. = ~70% of total feed energy requirements Calf-to-Weaning Calf-to-Beef

Traditionally, expressed Selection for FCR Feed efficiency FCR = feed intake : weight gain Similar to selection for growth rate Improves FCR in growing animal BUT Increase in cow size Increase in maintenance requirements Increase costs & Environment Improve the efficiency of growth / finishing phase BUT not necessarily the entire production system

France: Increasing cow size? Frame size of beef cows increased during the last 25 years genetics + feeding e.g. carcass weight of cull cows increased from 357 to 408 kg (Lherm et al., 2004). UK: Avg. cow live weight.considerably higher than 20 years ago, probably as a result of genetic selection for higher body weight 680 kg vs. 434 560 kg (Hyslop, 2006)

Residual Feed Intake (RFI) Alternative measure of feed efficiency independent of growth & body size Expected requirements for maintenance & growth RFI = animals actual intake - predicted intake Assume: 2 steers: LW = 600 kg: ADG = 1..0 kg EXPECT them to eat 10 kg DM/day BUT one eats 11 kg DM & one eats 9 kg DM Steer eating 1 kg more than expected = RFI (+1) = High = less efficient Steer eating 1 kg less than expected = RFI (-1) = Low = more efficient Therefore, negative or lower RFI values are better

Residual Feed Intake F e ed I ntake (U F V /d a y ) 12 10 8 6 4 High RFI = inefficient Low RFI = efficient 400 450 500 550 600 650 700 Live weight

In any population of cattle, within breed, = >LARGE variation in Feed Efficiency Table 1: Intake and performance of beef cattle with high (Inefficient), medium and low (efficient) RFI RFI 1 RFI 2 RFI 3 High Med. Low Sig. High Med. Low Sig. High Med. Low Sig. DMI (kg/day) 6.3 5.9 5.5 *** 10.2 9.6 9.0 *** 7.3 6.8 6.2 *** Live weight (kg) 316 327 330 NS 522 523 512 NS 254 255 252 NS ADG (kg) 0.60 0.61 0.60 NS 1.66 1.63 1.53 NS 1.52 1.49 1.54 NS Source: 1 Lawrence et al., 2012; 2 Fitzsimons et al., 2014; 3 Kelly et al., 2010 Top 1/3 vs Bottom 1/3 15% 13% 18% Scope: Breeding more feed efficient cattle!!

Factors affecting Feed Efficiency

Live weight & live weight gain Table 2: Theoretical energy requirements of finishing bulls (UFV/day) at different weights and growth rates Average daily gain (kg) Maintenance Live weight (kg) 450 500 550 600 650 0.0 4.4 4.8 5.1 5.5 5.8 1.0 6.0 6.5 6.9 7.4 7.9 1.2 6.5 7.0 7.5 8.0 8.6 1.4 7.0 7.6 8.1 8.7 9.4 1.2 vs. 1.4 0.5 0.6 0.6 0.7 0.8 Source: INRA Feed efficiency is better with light, fast growing animals

Duration of finishing

Daily gain (kg) by finishing Interval Interval (d) Silage + 6 kg Conc. 0 147 0.65 1.10 0 56 0.80 1.43 56 98 0.59 1.02 98 147 0.54 (-33%) 0.79 (-45%) Rate of live weight gain is NOT constant over the finishing period Initial increase in gut fill, Fixed/declining DMI relative to increased weight (& Reqs) Increased fat deposition (much less efficient)

Effect of Feeding Period on live/carcass weight gain & FCR Charolais xbred steers: High concentrate diet Finishing period 0-12 wks 12-23 wks Live weight gain (kg/d) 1.42 1.16 Carcass gain (kg/d) 1.04 0.84-19% Concentrate intake (kg DM/d) 10.2 11.4 +12% Feed efficiency (Conc DMI:gain) Live weight 7.2 9.9 Carcass 9.9 13.6-38%

Guidelines for duration of Finishing Period on ad libitum concentrate Days ADG (kg) Heifers 60-80 1.10-1.30 Steers 70-90 1.25-1.45 Bulls <180 (80-120) 1.70-1.90

Duration of Finishing Period Avoiding excessively long finishing periods & minimising carcass fatness at slaughter (without impairing carcass value) are ways to reduce feed requirements & feed costs associated with finishing cattle.

Breeds: Beef Dairy vs. Beef 20 Residual Feed Intake (MJ/d) Finishing period: 15 10 14% AN HE SI Compared to dairy breeds, beef breeds had: +23% LWG per unit of energy consumed 5 CH L BUT with higher KO% & greater proportion of meat in carcass 0-5 +51% meat per unit of energy consumed Source: Crowley et al. 2010 Source: Clarke et al. 2009

GENDER: Bulls vs. comparable steers Grange data: ~ +8% ADG ~ +9% carcass wt ~ +20% lean meat yield NI data:: (/kg feed eaten) ~ +10% ADG ~ +14% carcass wt ~ +20% lean meat ~ +17% saleable meat USA data: ~ +17% ADG ~ -35% fat ~ +13% feed efficiency Europe data: ~ +1% higher intake ~ +20% ADG ~ -20% fat ~ +17% feed efficiency

Indoor Finishing

Grass silage Grass silage - primary forage for feeding cattle over winter. Digestibility (DMD): primary factor influencing the nutritive value of grass silage & consequently, the performance of cattle offered grass silage-based diets. Both intake & animal performance increases with increasing digestibility.

Silage Digestibility - what difference does it make? DMD % 75 70 65 60 Harvest 20 May 2 June 15 June 30 June Date Silage DMI 9.0 8.3 7.6 7.0 Kg/day Animal gain (kg/day) Live weight 0.83 0.66 0.49 0.31 Carcass 0.51 0.39 0.27 0.15 Source: Teagasc, Grange

Effect of silage digestibility on carcass gain Daily carcass gain (g) / 10 g/kg increase in silage DOM 35 30 25 20 15 10 5 0 GS GS + CO. GS + CO 0.2-0.37 0.2-0.4 Steen, 1988 Steen et al. 2002

Response in carcass gain to a 1% unit change in grass silage digestibility at various forage:concentrate ratios 40 35 30 g/day 25 20 15 10 5 0 100:0 80:20 60:40 40:60 Keady et al. 2013

Effect of silage digestibility & concentrate level on carcass gain 30 25 Daily carcass gain (g) / 10 g/kg increase in silage DOM 20 15 10 5 0 2.25 4.5 6.75 9 Concentrate (kg/day) Steen, 1998

Implication Silage Digestibility Low DMD grass silage means that higher levels of supplementation are needed to maintain performance Each 1-unit decline in digestibility requires an additional ~0.4 kg concentrate daily to sustain performance in finishing cattle (Keady et al., 2013) At high levels of concentrate feeding, silage digestibility had no effect on carcass gain

Silage Intake & Substitution rate

Intake & concentrate supplementation 10 8 DM Intake (kg/day) 6 4 2 0 Supplementation increases total DMI, although at a progressively decreasing rate Silage DMI Total DMI 0 1 2 3 4 5 6 7 8 9 10 Concentrate (kg/day) Source: Teagasc, Grange

Substitution Rate - High Digestibility Silage 0.7 0.6 0.5 Kg silage DM / kg Conc. DM 0.4 0.3 0.2 0.1 0 0.20 0.25 0.30 0.35 0.40 0.45 0.50 Dietary concentrate proportion Steen; 1998; Agnew & Carson, 2000; Steen & Kilpatrick, 2000; Patterson et al., 2000; Dawson et al., 2002; Caplis et al. 2005

Substitution Rate - High Digestibility Silage 1.4 1.2 kg silage DM / kg conc. DM 1 0.8 0.6 0.4 0.2 0 0.2 0.3 0.4 0.5 0.6 0.7 0.8 Dietary concentrate proportion Steen, 1998; Caplis, 2004

Effect of silage digestibility on substitution rate 1.2 1 kg silage DM / kg conc. DM 0.8 0.6 0.4 0.2 0 Low D High D. Low D High D Drennan & Keane, 1987 Steen, 1998

Effect of energy supplement type Finishing Cattle on substitution rate Starch = Fibre = Sugar (Moloney et al. 1993) Starch = Fibre (Steen, 1995; O Kiely & Moloney, 1994) Starch < Fibre (Moloney, 1996) Starch < Fat (Steen, 1995; Moloney, 1996)

Effect of dietary concentrate proportion 0.85 on diet digestibility D igestibility 0.8 0.75 0.7 0.65 0.6 Implication: Dietary energy intake often mirrors DMI 0 0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 Concentrate proportion Steen & Robson, 1995; Steen & Kilpatrick, 2000; Patterson et al., 2000; Caplis, 2004

Effect of dietary concentrate proportion on fibre digestibility D igestibility 0.9 0.85 0.8 0.75 0.7 0.65 0.6 0.55 0.5 0.45 0.4 CF Negative associative effect NDF CF ADF P<0.001 0 0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 Concentrate proportion Steen & Robson, 1995; Steen & Kilpatrick, 2000; Patterson et al., 2000; Steen et al. 2002

Production response to concentrate supplementation

Concentrate supplementation - finishing cattle 300 250 Days to finish 200 150 100 50 0 0 0.3 0.7 Dietary concentrate proportion Scollan et al. 2003

Effect of concentrate supplementation on growth response - steers Liveweight gain (kg/day) 1.4 1.2 1 0.8 0.6 0.4 0.2 0 Incremental growth response declines as concentrate level increases Higher response in animals with high growth potential e.g. gender, breed, compensatory growth 0 1 2 3 4 5 6 7 8 9 10 Bull Steer Concentrates (kg/day) Source: Teagasc, Grange

Effect of silage digestibility on carcass growth response to supplementation Carcass gain (kg/day) 0.9 0.8 0.7 0.6 0.5 0.4 0.3 0.2 0.1 0 High DOM Low DOM 0 1 2 3 4 5 6 7 8 9 Concentrates (kg/day) Steen, 1998

g /d a y Concentrate supplementation & 1800 1600 1400 1200 1000 800 600 400 200 0 Growth response 0.2 0.3 0.4 0.5 0.6 0.7 0.8 Lwg Carcass Lean O = 0.95 Dietary Concentrate Proportion Patterson et al. 2000

Optimum level of concentrate feeding? Table 4: Cost (c/kg) of carcass gain for steers Feeding level Concentrate costs ( /t) (kg/day concentrate) 175 200 225 250 275 300 3 117 147 178 209 239 270 4 119 155 190 225 261 296 5 124 165 207 249 290 332 6 159 214 268 323 378 433 7 172 242 312 382 452 522 8 180 280 380 480 580 680 Silage substituted valued at 15.0 cents/kg DM

Ad libitum feeding of concentrates for finishing cattle Considered when Silage digestibility is low Animal growth potential is high Especially where silage supplies are in deficit Critical to ensure: 1. gradual adaptation to the concentrates, 2. minimum roughage inclusion (~10% of total DM intake) in the diet for rumen function, 3. that meal supply never runs out 4. that a constant supply of fresh water is provided.

Ingredient Composition

Concentrate Energy source & Growth response Supplements to grass silage Wheat = Barley (Steen, 1993; Drennan et al. 2006) Sugar-based = Starch-based @~0.2 DMI Starch-based > or < Fibre-based (Drennan, 1985; Moloney et al. 1993; Chapple et al. 1996) (Moloney et al. 1993; O Kiely & Moloney, 1994; Steen, 1995; Moloney, 1996; Moloney et al. 2001) Fat-based < Starch or Fibre-based (Steen, 1995; Moloney, 1996)

Concentrate type 4 rations x 2 feeding levels Ration A. Rapidly fermentable starch - RFS (Barley-based) B. Slowly fermentable starch SFS (Maize-based) C. RFS + digestible fibre - RFS+F D. Digestible fibre-based - Fibre (Pulp) Concentrate feeding level Ad libitum 5 kg per head/day Formulated: ~ same Energy & Protein concentration

Concentrate type Ration Type RFS SFS RFS+F Fibre Sig. Intake (kg DM) Silage 3.8 4.0 3.8 3.6 ns Conc. 6.5 6.5 6.9 6.5 ns Total 10.2 10.5 10.7 10.1 ns Daily gain (g) Live weight 1043 985 1032 911 ns Carcass 582 570 584 520 ns KO (g/kg) 537 541 540 539 ns Fat 3.5 3.4 3.5 3.5 ns FCE 56.8 54.6 55.6 52.0 ns g carcass / kg DMI McGee et al. 2006, 2009

Concentrate type: Maize meal Dairy bulls 170 days Ration Type Barley-based Barley/Maize-based Sig. Intake (kg DM) 9.21 9.54 * Live weight gain (g/day) 1697 1745 ns Carcass wt (kg) 275 279 ns Carcass Fat (1-5) 2.98 3.09 ns Keane, 2008

Response to protein in Finishing Cattle High concentrate diet Gender Bulls Heifers Protein + - + - Start wt (kg) 331 332 288 286 LWG (kg) 1.58 1.55 1.25 1.22 Slaughter wt (kg) 572 568 470 463 Carcass wt (kg) 319 315 255 253 Kill-out (g/kg) 558 555 542 547 Intake (kg DM/d) 9.4 9.1 8.0 7.7

Composition (g/kg) of diets for bulls & heifers Protein + - Rolled barley 875 945 Soya-bean meal 70 - Molasses 40 40 Mins/vits 15 15 CP (g/kg DM) 143 116

Response to protein in Finishing Cattle Grass silage + concentrates Finishing steers / heifers / bulls Barley-based conc. + Protein: High DMD silage = X Low DMD silage = Low crude protein grass silage = Implications With low DMD & low CP grass silage % CP in Dietary DM Bulls: growing 13-15 Bulls: Finishing 12-13 Heifers/Steers 11-12 Higher CP % required in concentrate [~11-12% to ~14-20%, depending]

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