How Fiber Digestibility Affects Forage Quality and Milk Production

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How Fiber Digestibility Affects Forage Quality and Milk Production Dr. Dave Combs Professor Dept of Dairy Science University of Wisconsin-Madison What causes performance swings in dairy diets? MOST OF THE TIME ENERGY üdiet Energy is impacted largely by carbohydrates ü Fiber ü Starch ü Fiber is always lower energy than starch (grain) ü 2-3 unit drop in Fiber or Starch digestibility will decrease milk by about one pound New Technologies and Innovations in Forage Feeding Programs for Livestock Digestibility! Corn Silage Shredlage ( starch digestibility) BMR ( fiber digestibility) Alfalfa Reduced lignin ( NDF digestibility) Grasses Improved grasses for high producing dairy cows (Higher fiber digestibility than alfalfa or corn silage) Forage testing/analysis Indigestible fiber (undf 240 ) Total Tract NDF digestibility () Topic #1. What makes a better forage? ühigh digestibility ü Fiber (-) ü Fiber digestibility (+) ühigh intake potential ü Fiber (-) ü Fiber digestibility (+) BOTH NDF and NDF digestibility are needed to assess forage quality Forage Fiber Tests Fiber digestibility varies in forages Test Rumen Fill TDN Estimation Diet Formulation Herd Diagnostics NDFOM X X X X Quality Index NDFD(30 or 48) X/? X X/? X X X X X undf240 X? NDF kd RFV/RFQ Milk/ton X X X Range in % of NDF Alfalfa hay and silage 25-70 Corn silage 25-80 Grass hay and silage 15-80 Two units increase in diet can potentially increase milk yield by 1 lb Combs University of Wisconsin-Madison 1 of 6

How variable is alfalfa fiber digestibility? + 2 TDN + 1 SD Mean Why is fiber digestibility important? Oba and Allen (1999) A 1% change in vitro or in situ NDF digestibility (primarily 30-h or 48-h NDFD) was correlated with: ü 0.4 lb increase in dry matter intake ü 0.5 lb increase in 4% fat corrected milk yield 1/3 of alfalfa forages at the same NDF will vary by more than 2 units of TDN from average - 1 SD - 2 TDN 1: Maturity NDF Lignin % of DM % of DM % of NDF Immature 33 5.4 54 Vegetative 37 6.2 50 Mid-maturity 43 7.3 47 Mature 50 8.4 46 2: Growing conditions/environment ümoisture ütemperature üsun intensity 2/3 or more of variation in fiber digestibility is likely due to growing conditions/environment 3: Genetics: Reduced Lignin Alfalfa 2: Genetics HiGest TM Alforex HarvXtra TM Forage Genetics International Variety HiGest TM (Alforex) HarvXtra TM (FGI) Lignin Reduction Unit reduction (assuming 7% lignin) 7 to 10% 0.49 to 0.7 10 to 15% 0.7 to 1.05 Combs University of Wisconsin-Madison 2 of 6

Effect of harvest delay on NDFD, average of 4 cuttings (HarvExtra Alfalfa) Effect of Low Lignin Trait on Alfalfa ADF and NDF Digestion NDFD, % of NDF 53 51 49 47 45 43 41 39 37 35 7 day delay in quality decline 28 32 35 Growth cycle Days RR RR/RL Undersander, 2015 Roundup Harvest Ready interval Roundup Ready + Low P Value Item Lignin* Forage ADF 28d 26.6 26.5 NS NS 33d 27.2 26.6 35d 27.8 26.5 Harvest interval 28d 52.1 56.3 0.05 0.01 33d 46.3 51.9 35d 46.8 51.1 * HARVXTRA Forage Genetics International Li, Li, Undersander and Combs, 2015, ADSA abstract + 8% +12% + 9% Nutrient composition of corn silage stalk hybrids Item BMR CONS HFD LFY SEM P-value DM, % as fed 33.7 34.5 35.1 33.2 0.9 0.45 CP, %DM 8.0 7.8 8.1 8.0 0.2 0.20 NDF, %DM 42.3 42.6 45.0 42.3 0.8 0.09 Lignin, %DM Starch, %DM 2.0 b 2.8 a 2.9 a 2.6 a 0.2 0.001 28.7 ab 30.1 a 26.7 b 30.0 ab 1.1 0.02 DMI & Milk Yield greater in BMR/HFD Item BMR CONS HFD LFY SEM P-value DMI, kg/d 25.1 a 24.0 b 24.6 a 23.0 b 0.5 0.001 Milk, kg/d 38.6 a 37.2 b 38.1 a 37.4 b 0.8 0.001 Fat, % 3.55 3.62 3.61 3.64 0.08 0.25 kg milk/kg DMI 1.52 1.54 1.55 1.55 0.03 0.61 Ferraretto and Shaver, 2013 4: Harvest management ümoisture (leaf shatter) ürain damage ürespiration losses due to slow dry-down Fiber in leaves is higher in digestibility than fiber in stems Using forage analysis to assess quality Reading Forage Analyses ü Dave s Quick List: 1. NDF and Starch 2. Protein 3. Ash Content ü Evaluate Digestibility ü ü StarchD Combs University of Wisconsin-Madison 3 of 6

TOPIC # 2: Assessing fiber digestion Forage Quality Indicators for High-Producing Dairy Herds Parameter NDF Lignin Indicates Better Quality Primary Reason Rumen Fill Limitation of DMI undf 240 NDFD 30 Potential for production response or feeding of higher-forage diets Energy Density Poor digestion < 40% Excellent digestion > 50% A 2-3 unit change in fiber digestibility corresponds to 1 lb change in milk yield. NFC CP Ash RFV; RFQ Minimal Soil Contamination Potential for production response or feeding less corn grain Supplemental Protein Energy Density Quality Index for Ranking How Can We Equate Feed Fiber Measurements to Animal Utilization of NDF The Process of fiber digestion is described with the assay Feed and cow factors both affect fiber digestion? Feed fiber (NDF) Potentially digestible NDF Rate of fiber digestion Fiber digestion Rate of passage Fiber digestibility () Fiber digestion is affected by: How is determined? Feed characteristics üthe amount of fiber (NDF) ü Potentially digestible fiber (pdndf) (pdndf = NDF-uNDF 240 ) ü Rate of fiber digestion (kd) Animal and diet ü Intake affects rate of fiber passage (kp) ü Approx. 90% of NDF digestion is in rumen Forage sample Rate of fiber digestion (kd) Potentially digestible NDF (pdndf) Rate of fiber passage, (kp) Standardized iv NDFD (24, 30, 48h) and indf Rumen and hindgut digestion PD NDF * kd/(kd +kp) (total tract NDF Digestibility) Combs University of Wisconsin-Madison 4 of 6

combines in vitro rate of NDF digestion with indf to improve the prediction of in vivo fiber digestion Feed Analysis Lab Report can be quickly and cheaply analyzed by NIRS is a prediction of NDF digestibility for a feed (or diet) in 650 kg cow consuming 24 kg DM of a 28-30% NDF diet. 47.98 42.34 Lopes et. al. 2015 JDS Low NDF corn silage (good) Which is the better Haylage? BUT- Lower than average fiber digestibility (bad) Typical NDF and values NDF Average High Low Alfalfa < 40% 42% > 48% < 36% Corn Silage < 40% 42% > 48% < 36% Grasses < 45% 42% > 48% < 36% Dairy quality alfalfa and corn silages will be < 40% NDF with a value of at least 42% A 2-3 unit change in ration corresponds to a one pound change in milk yield. Milk predictions from a model based on NRC equations and as the coefficient for NDF digestibility Predicted milk yield, kg 50.0 45.0 40.0 35.0 30.0 25.0 25.0 30.0 35.0 40.0 45.0 50.0 Observed milk yield, kg Combs University of Wisconsin-Madison 5 of 6

The Take Home Message 1. Fiber digestibility has a big impact on milk yield. A 2-3 unit change in ration corresponds to a one pound change in milk yield. Thank You! Visit our Web sites: http://dysci.wisc.edu/ http://fyi.uwex.edu/forage/ 2. The test was developed to predict fiber digestibility in high producing dairy cattle Can be used across forage types and byproduct feeds Can be used in ration balancing and evaluation Combs University of Wisconsin-Madison 6 of 6