A pre-publication draft: In R. Burgess & K. MacDonald (Eds.) (2005). Evolutionary perspectives on human development. CA: Sage Publications

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1 Evolutionary reconceptualization of Kohlberg 1 A pre-publication draft: In R. Burgess & K. MacDonald (Eds.) (2005). Evolutionary perspectives on human development. CA: Sage Publications An Evolutionary Reconceptualization of Kohlberg s Model of Moral Development Dennis Krebs Simon Fraser University

2 Evolutionary reconceptualization of Kohlberg 2 An Evolutionary Reconceptualization of Kohlberg s Model of Moral Development At the end of a program of research my colleagues and I conducted on Kohlberg s (1984) model of moral development, I concluded that it is poorly equipped to account for the kinds of moral judgments and moral behaviors people emit in their everyday lives. My search for a more ecologically valid model of morality led me to what many people would consider an unlikely source, the modern synthetic theory of evolution. In this chapter I will briefly describe Kohlberg s model of moral development, identify its deficiencies, reconceptualize Kohlbergian structures of moral judgment as evolved decision-making mechanisms, and explain why I believe this reconceptualization gives rise to a more valid model of moral development. Kohlberg s Model of Moral Development The two central assumptions of Kohlberg s (1984) model of moral development are that the primary source of morality is moral reasoning and that people become more moral because they acquire increasingly sophisticated structures of moral reasoning. To assess moral reasoning, Kohlberg and his colleagues developed a test containing nine hypothetical moral dilemmas followed by sets of questions (Colby & Kohlberg, 1987). In the most often cited dilemma, a character named Heinz must decide whether or not to steal an over-priced drug to save his dying wife. Kohlbergians assume that the deontic decisions people make whether Heinz should or should not steal the drug stem from structures of moral reasoning. To map these structures, Kohlbergians ask moral decision-makers why questions designed to induce them to explicate the principles from which they deduced their moral choices. The main source of support for Kohlberg s model stems from a longitudinal study conducted by Kohlberg and his colleagues. In 1958 Kohlberg gave a cohort of boys an early version of his test, and with the help of colleagues, retested them every three or four years for

3 Evolutionary reconceptualization of Kohlberg 3 more than two decades (Colby & Kohlberg, 1987). This study produced three main conclusions. First, moral judgments made by individuals are structurally consistent across varying content. That is to say, individuals invoke the same forms of moral reasoning to resolve different kinds of moral dilemma; moral judgment is organized in structures of the whole. Second, as children grow older, their structures of moral reasoning normally undergo qualitative changes. According to Colby and Kohlberg (1987), new structures transform and displace older structures, giving rise to the five stages of moral development summarized in Table 1. Kohlberg and his colleagues found that the participants in his longitudinal study went through these stages in an invariant sequence. Third, because the structures of moral reasoning that define higher stages of moral development are more cognitively sophisticated than those that define lower stages that is to say, more differentiated, integrated, and logical and based in more sophisticated perspectivetaking abilities, they give rise to more moral decisions (that is to say, moral decisions that are more prescriptive, universal, impartial, and objective) Insert Table 1 about here In 1984, Kohlberg and Candee published a model linking moral reasoning to moral conduct. Kohlberg and Candee (1984) argued that moral judgment is necessary but not sufficient for moral behavior. To behave morally, people must decide what the most moral course of action is, but this does not necessarily compel them to behave morally. To implement moral decisions, people must experience an obligation to carry them out, which entails making followup judgments of responsibility. In addition, people must possess sufficient ego-strength to do what they think they should.

4 Evolutionary reconceptualization of Kohlberg 4 An Evaluation of Kohlberg s Model of Moral Development Several years ago, my colleagues and I noted that most of the empirical support for Kohlberg s model stemmed from studies conducted in academic contexts in which trained interviewers or experimenters asked university students to make and to justify decisions about how the characters in Kohlberg s dilemmas should resolve hypothetical moral conflicts. We set out to vary parameters of this prototypic situation and determine whether they affected the forms of moral judgment people invoked. To this end, we asked people who differed in a variety of ways to make moral judgments in a variety of non-academic contexts, to audiences other than experimenters, about moral dilemmas other than those on Kohlberg test, and about people other than the characters in Kohlbergian dilemmas. The findings from this program of research are reviewed in several publications (see Krebs, Denton & Wark, 1997; Krebs, Vermeulen, Carpendale & Denton, 1991; Krebs, Wark & Krebs, 1995, and Wark & Krebs, 1997). To summarize, we found, as Kohlberg would expect, that people tend to make structurally similar that is to say same-stage moral judgments to hypothetical moral dilemmas similar to those on Kohlberg s test. However, we also found that people tend to make lower-stage moral judgments about more real-life-like dilemmas. For example, compared to moral judgments on Kohlberg s test, participants in our studies made relatively low-stage moral judgments to moral dilemmas about prosocial behavior and drinking and driving (Krebs, Denton, Vermeulen, Carpendale, & Bush, 1991), business deals and free trade (Carpendale & Krebs, 1995), prostitution (Bartek, Krebs & Taylor, 1993), and a variety of real-life dilemmas (Krebs, Denton, Wark, Couch, Racine, & Krebs, 2002; Wark & Krebs, 1997). Young adults made lower-stage moral judgments about Kohlbergian dilemmas and a dilemma about drinking and driving when they were drinking at bars and parties than they did in academic

5 Evolutionary reconceptualization of Kohlberg 5 contexts (Denton & Krebs, 1990). University students made lower-stage moral judgments to a Professor of Business Administration than to a Professor of Philosophy (Carpendale & Krebs, 1992). Parents and children made relatively low-stage moral judgments about moral dilemmas they experienced in their families (Krebs, Vermeulen & Denton, 1991). Incarcerated female juvenile delinquents, especially those who scored high on a test of defensiveness, made relatively low-stage moral judgments about moral dilemmas involving prostitution (Bartek, Krebs & Taylor, 1993). People tended to make different kinds of moral judgments about themselves than they did about others (Krebs et al., 2002; Krebs & Laird, 1998). Considered as a whole, the findings from our research program did not support Kohlberg s model of moral development. Our data were not consistent with the assumption that moral judgments stem from structures of the whole that have transformed and displaced their predecessors. Instead, our data suggested that people retain old structures of moral judgment after they develop new ones. Moral development is characterized more by an expansion of the range of possibilities than by changes in the overriding structure of the mind. The types of moral judgment people make are not determined solely by the structures of moral reasoning they have acquired; moral judgments are the product of complex interactions between internal qualities of people and the external situational variables they encounter. Findings from our studies also suggested that the relation between moral judgment and moral behavior is more complex than Kohlberg and Candee (1984) assumed. As postulated by Kohlberg and Candee, in some contexts people appear to decide what is right, then act on their decisions. However, in other contexts people appear to act without engaging in moral reasoning, then invoke moral judgments to justify their decisions (Denton & Krebs, 1990; Krebs et al., 2002; Krebs & Laird, 1998).

6 Evolutionary reconceptualization of Kohlberg 6 In the end I concluded that Kohlberg s model pertains to only one of many functions of moral judgment, namely to make the most rational moral decision of which one is capable. The methods employed by Kohlbergians are designed to induce people to perform this function. Making moral judgments about Kohlbergian dilemmas in academic contexts is akin to solving problems in logic. As acknowledged by Colby and Kohlberg (1987), Kohlberg s model is a model of moral competence. To adequately account for moral development, we must attend to other functions of moral judgment and moral behavior. In real-life people use moral judgments for more social and pragmatic purposes than they do when they make moral judgments on Kohlberg s test. When people tell other people they should behave in particular ways, they are rarely trying to explicate their ideal conceptions of morality; rather, they are usually trying to induce the recipients of such judgments to behave in accordance with the moral prescriptions they are pronouncing. They are trying to exert social influence. Recipients of such moral judgments often respond by arguing with those who pronounce them, attempting to persuade them that the prescriptions are unjustified. Negotiations ensue in which parties adduce arguments to support their points of view (Krebs et al., 2002). In real life, people also use moral judgments to approve and disapprove of the behavior of others, to praise, condemn, and blame. In addition, people use moral judgments to guide, justify, and excuse their own behavior to approve and disapprove of themselves, to enhance their social image and to foster self-esteem (Denton & Krebs, 1990; Krebs et al., 2002; Krebs & Laird, 1998). A Functional Reconceptualization of Kohlberg s Elements of Morality As I have explained, when people make decisions about what the characters in Kohlbergian dilemmas should do, interviewers ask them why, and after interviewees proffer a

7 Evolutionary reconceptualization of Kohlberg 7 reason, interviewers ask them why again until interviewees reach the limit of their explanations. For example, an interviewee might say, Heinz should steal the drug to save his wife s life, and the interviewer might ask, why should Heinz save his wife s life? In response to this probe, the interviewee might answer, because he owes it to her. The interviewer might then ask, why is it right for Heinz to fulfill his obligations to his wife? and the person might say, because she has done a lot for him or because he made a marriage contract, and so on. Colby and Kohlberg (1987) have classified the reasons people adduce in support of their decisions and labeled them the elements of morality (see Table 2). From Kohlberg s perspective, why questions induce people to explicate the principles from which they derived their moral decisions. From a more functional perspective, such questions induce people to identify the goals they are advising moral decision-makers to pursue, goals such as doing their duty, paying their debts, reciprocating, obtaining approval, avoiding punishment, enhancing their reputations, supporting their groups and so on (see Table 2). From a Kohlbergian perspective, the purpose of repeatedly asking why questions is to induce people to invoke increasingly abstract principles of morality. From a functional perspective, the purpose of repeatedly asking why questions is to get at increasingly ultimate goals and purposes. If you keep asking people functional why questions, you will ultimately end up in one of two places, with God or in the theory of evolution. I ended up in the latter Insert Table 2 about here An Earlier Examination of Relations Between Kohlbergian and Evolutionary Approaches to Moral Development I was not the first psychologist to adopt a biological approach to morality and view

8 Evolutionary reconceptualization of Kohlberg 8 Kohlberg s model from an evolutionary perspective. Kevin MacDonald (1988) was. MacDonald (1988) reviewed evidence on moral judgment and moral behavior and concluded that much of it was more consistent with a sociobiological viewpoint than with Kohlberg s cognitivedevelopmental model. In particular, MacDonald (1988) adduced evidence supporting the following eight conclusions. First, children s moral behavior is more self-interested than it appears from the conclusions reached by researchers. Second, people invoke different forms of moral reasoning in different contexts. Third, people perform cost-benefit analyses when they make real-life moral decisions. Fourth, reasoning about oneself, one s relatives, and significant others is done with a different calculus than is reasoning about hypothetical situations (p. 148). Fifth, people sometimes fail to behave in accordance with their moral judgments, especially when they conflict with their self-interest. The link between moral reasoning and moral behavior is weak except when the decisions in question involve little cost. Sixth, people use moral judgments to create false impressions and to justify their behavior: Individuals who are adept at moral reasoning (i.e., perform at the higher stages) are better able to provide reasons which rationalize their self-interested actions in a manner that would justify their behavior to other individuals (p. 143). Seven, only the first three stages in Kohlberg s sequence are crossculturally universal. And finally, emotional reactions play an important role in real-life moral decision-making. In the remainder of this paper, I build upon these ideas. I review the basic propositions of evolutionary theory, reconceptualize the structures of moral judgment described by Kohlbergians as evolved mental mechanisms, offer an account of the evolution of moral judgment and moral behavior, and explain why I believe that this account gives rise to a more ecologically valid model of moral development than the one advanced by Kohlberg and his colleagues.

9 Evolutionary reconceptualization of Kohlberg 9 Toward an Evolutionary Model of Morality Evolutionary theorists assume that the ultimate goal of most (but not all) behavior is to enhance actors inclusive fitness, defined in terms of the number of replicas of their genes they contribute to future generations. More precisely, evolutionary theorists assume that individuals inherit genes that were selected in ancestral environments because they guided the development of mechanisms that enabled their ancestors to survive, to reproduce and to pass the genes and mechanisms on to future generations. The function of evolved mechanisms is to help individuals solve adaptive problems pertaining to survival, reproduction, care of offspring, and so on. There are two basic ways in which genes, or mechanisms, can be selected and increase in frequency, or evolve, over generations. In the first, the vehicles housing the genes or mechanisms that is to say, individuals survive until sexual maturity, pass replicas of their genes or mechanisms on to others through asexual or sexual reproduction, then take whatever measures are necessary to insure that the offspring in whom they have deposited replicas of their genes survive and reproduce. In the second, individuals behave in ways that foster the survival and reproduction of individuals other than their offspring who possess replicas of their genes that is to say, individuals who are similar to them genetically. This process is usually referred to as kin selection. The main difference between the two forms of natural selection is that in the first individuals create the vehicles that transport replicas of 50 % of their genes to future generations, whereas in the second individuals assist individuals created by others, who may possess varying proportions of their genes. Evolved Psychological Mechanisms Evolutionary theorists may attend to different products of evolution. For example, some

10 Evolutionary reconceptualization of Kohlberg 10 theorists attend to the frequency of genes in a population, whereas others attend to the number of offspring produced by individuals. Evolutionary psychologists attend to the mental mechanisms that individuals inherit and the behavioral strategies such mechanisms contain. Buss (1999, pp ) has identified six properties of evolved psychological mechanisms. First, they are shaped by the ways in which they recurrently solved specific adaptive problems over the evolutionary history of the species in which they evolved. Second, they are activated by relatively narrow slices of information. Third, the information that triggers such mechanisms the input pertains to the particular adaptive problems the mechanisms were designed to solve. Fourth, the information processed by such mechanisms is transformed by a set of if-then decision rules into output. Fifth, the output of an evolved psychological mechanism can be physiological activity, information to other psychological mechanisms, or manifest behavior. Finally, the output is directed toward the solution to a specific adaptive problem. Although it is safe to assume that the strategies implicit in such mechanisms were on average winning strategies in ancestral environments, they need not necessarily produce optimal solutions to the problems that activate them in modern environments. Kohlberg s Structures of Moral Judgment Reconceptualized as Evolved Psychological Mechanisms. In contrast to learning theorists who posit general purpose mechanisms, both Kohlbergians and evolutionary psychologists believe the mind is composed of specialized structures or mechanisms designed for particular purposes. Kohlbergians and evolutionary theorists assume that the mental structures that give rise to moral judgments and moral behaviors are activated by particular kinds of problem and that their function is to process information in ways that give rise to particular kinds of decision. Both types of theorist assume that the form or

11 Evolutionary reconceptualization of Kohlberg 11 pattern of the output produced by mental structures reflects the ways in which they are designed, and that the primary goal of social scientists is to map the design of the mechanisms by deciphering the operating principles or decision-rules implicit in their output. Kohlbergian and evolutionary psychologists also are attentive to the origin and development of mental structures and mechanisms. Such similarities notwithstanding, Kohlbergians make different assumptions from evolutionary theorists about how mental structures originate and the functions for which they were designed. Adopting an ontogenetic perspective, Kohlbergians assume that structures of moral judgment develop through the cognitive processes of assimilation and accommodation and that their function is to enable people to deduce the most moral solution to moral problems. Adopting a phylogenetic perspective, evolutionary theorists assume that evolved mechanisms originate from genetic variations selected in ancestral environments, and that their function is to help individuals solve real-life adaptive problems. Kohlbergians focus mainly on the cognitive functions of making moral judgments; evolutionary theorists focus mainly on the adaptive functions of moral behaviors. The central thesis of this paper is that the conception of mental mechanisms derived from evolutionary theory offers a better basis than Kohlberg s conception of structures of moral judgment for explaining the ways in which people make moral decisions and for accounting for moral development. Stated in stronger terms, I believe that most of the moral judgments people make in their everyday lives and most of the moral behaviors they emit are products of the kinds of evolved psychological mechanisms described by evolutionary psychologists, as opposed to the structures of moral reasoning described by Kohlbergians. The Evolution of Moral Mechanisms If we assume that people derive moral judgments from evolved mental mechanisms, the

12 Evolutionary reconceptualization of Kohlberg 12 first question that arises is, what adaptive functions did such mechanisms serve in ancestral environments; what kinds of adaptive problems did such mechanisms help our hominid ancestors solve? I believe the answer to this question is that the mechanisms that give rise to morality evolved to solve the social problems that occurred when our ancestors banded together to foster their biological interests. Among the many species in the animal kingdom, Homo Sapiens are among the most social. From the perspective of evolution, mechanisms that dispose animals to aggregate and interact with other members of their species and for that matter with members of other species evolve when the mechanisms help animals foster their biological interests. Such mechanisms may help animals enhance their inclusive fitness in several ways. As examples, individuals who band together may be less susceptible than more solitary individuals to predators, and groups may be able to hunt larger game than individuals could kill on their own. Many of the benefits of sociality stem from cooperative exchanges. Individuals working together can often obtain more for themselves than they could by working alone. Individuals can often trade items of relatively little value for items of greater value. Individuals can render valuable assistance to others at little cost to themselves in return for low-cost assistance when they are in need. In such exchanges both (or all) parties can come out ahead through what economists call gains in trade. According to most experts on human evolution, cooperation was instrumental in the evolution of the human species. To quote Leakey and Lewin (1977), Throughout our recent evolutionary history, particularly since the rise of a hunting way of life, there must have been extreme selective pressure in favor of our ability to cooperate as a group The degree of selective pressure toward cooperation was so strong, and the period over which it operated so extended, that it can hardly have failed to have become embedded to some degree in our genetic

13 Evolutionary reconceptualization of Kohlberg 13 makeup (p. 45). In view of the benefits of cooperation, it might seem that mechanisms disposing individuals to cooperate should evolve without impediment, but unfortunately this is not the case. In his book, Theory of Justice, the philosopher John Rawls (1971) explains why: Although a society is a cooperative venture for mutual advantage, it is typically marked by a conflict as well as by an identity of interests. There is an identity of interests since social cooperation makes possible a better life for all than any would have if each were to live solely by his own efforts. There is a conflict of interests since persons are not indifferent as to how the greater benefits of their collaboration are distributed, for in order to pursue their ends, each prefers a larger to a lesser share. (p. 4) Selfish preferences pose a serious problem for the evolution of cooperative mechanisms, because it is in everyone s biological interest to give less than his or her share and to induce others to give more than their share. Inasmuch as the resources in question foster the inclusive fitness of those who possess them, individuals who inherit mechanisms that dispose them to behave selfishly could fare better than those who inherit mechanisms that dispose them to behave cooperatively, mediating the evolution of selfishness and the extinction of cooperation. However, fortunately for the evolution of morality, it is not in individuals interest to let others treat them selfishly, so we would expect mechanisms designed to counteract selfishness in others to evolve. Most people consider selfish behaviors, defined as fostering one s own interests at the expense of others, immoral; and most people consider cooperative behaviors, defined as fostering one s interests in ways that foster the interests of others, moral. I believe that the biological function of morality is to uphold fitness-enhancing systems of cooperation. Viewing morality in these terms helps us understand its nature. Morality boils down to giving one s share (doing one s duties) and taking one s share (exercising one s rights), cooperating with others by treating

14 Evolutionary reconceptualization of Kohlberg 14 them fairly, as one would like to be treated, and resisting the temptation to maximize one s gains at the expense of others. This conception of morality has been espoused in non-biological terms by many scholars. As one example, Rest (1983) asserts that morality consists in standards or guidelines that govern human cooperation in particular how rights, duties, and benefits are [to be] allocated Moralities are proposals for a system of mutual coordination of activities and cooperation among people (p. 558). The Social Functions of Morality In contrast to the purpose of making moral judgments about the hypothetical characters in Kohlbergian dilemmas, people make moral judgments in their everyday lives for two primary purposes: (a) to influence the behavior of others and (b) to guide their own behavior. Let us consider each in turn. Moral Judgments as Forms of Social Influence Viewed biologically, moral judgments are a form of communication. Biological analyses of communication assume that animals are evolved to send signals that induce recipients to behave in ways that foster the senders interests, or to manipulate them. Such signals are often deceptive (Dawkins, 1989; Mitchell & Thompson, 1986). Humans relatively large brains and their capacity for language enable them to employ a significantly larger range of manipulative communication strategies than those available to other species. Senders are able to take the perspective of recipients (referred to as mind-reading by some psychologists), and plan long into the future. Recipients reactions to senders signals are less a function of the physical properties of the signals themselves and more a function of how recipients represent them mentally. From a biological perspective, when people send second-person moral judgments to

15 Evolutionary reconceptualization of Kohlberg 15 others such as, you should help me, you should keep your promises, and you are not being fair to me, they are tying to induce recipients to foster their interests by persuading them to behave in accordance with the prescriptions. When people express more abstract judgments such as honesty is the best policy and people should obey the law, they are attempting to induce recipients to uphold the systems of cooperation from which they benefit. When senders support such judgments with reasons such as, because you will cultivate a good reputation and everyone will benefit in the end, they are attempting to induce recipients to form cognitive representations of the if conditions that activate the behaviors they are prescribing. Such judgments are tools of social influence, tactics of persuasion designed to induce recipients to behave in ways that directly or indirectly benefit those making the judgments. The natural selection of moral judgments. Of all the moral judgments people could make, why do they make those observed and classified by Kohlberg and his colleagues? Why do only a few of the many possible moral judgments become normative in all cultures? Kohlbergian and evolutionary psychologists would agree that people select the moral judgments that are best equipped to solve the moral problems they face, that such problems are social in nature, and that they involve conflicts of interest. However, Kohlbergians would focus on ideal solutions to hypothetical versions of such problems, whereas evolutionary theorists would focus on actual solutions to real problems in which the individuals involved are attempting to foster their biological interests. Although it might seem that individuals motivated to foster their biological interests would make moral judgments such as, you should sacrifice your interests for my sake, such judgments would not be effective, because recipients would not be receptive to them. In effect recipients of moral judgments are agents of selection, determining what kinds of judgment work.

16 Evolutionary reconceptualization of Kohlberg 16 Although those who make moral judgments may well attempt to use them to manipulate and exploit recipients, successful manipulations would have to be subtle, leading recipients to believe that abiding by them would foster their interests. In general, the most effective moral judgments should prescribe behaviors that foster the interests of senders and recipients. Thus, I would expect moral judgments to assume the form, you should foster my interests in ways that foster your interests, which is exactly what most of the moral judgments classified by Kohlbergians do. To summarize, senders do not exhort recipients to behave selfishly because it is not in senders interest, and senders do not exhort recipients to behave altruistically because recipients are unreceptive to such exhortations. Senders exhort recipients to adopt the kinds of cooperative strategies prescribed by Kohlbergian moral judgments because the mental mechanisms possessed by recipients are designed in ways that make them receptive to such judgments. The activation of stage-based moral judgments. As discussed, we found that in certain contexts adults invoke moral judgments made by people at all of the stages described by Kohlberg. From an evolutionary perspective, we would expect people to make the kinds of moral judgment that contain the greatest potential to foster their biological interests, which will depend on the if conditions implicit in the problems they encounter. So, for example, relatively powerful members of groups should make Stage 1 judgments that exhort weaker members to obey authority in order to avoid punishment. Those who are relatively equal in power should be more inclined to make Stage 2 judgments to each other that uphold mutually beneficial deals. Friends and relatives should make Stage 3 moral judgments that uphold their long-term relationships; and those who have a vested interest in maintaining the social order should make Stage 4 judgments. In general, the benefits implicitly promised to those who conform to high-stage moral judgments are more general and delayed than the benefits implicitly

17 Evolutionary reconceptualization of Kohlberg 17 promised to those who conform to low-stage moral judgments. Therefore, in general, the if conditions invoked to activate higher-stage moral behavior are more tenuous than the if conditions invoked to activate lower-stage moral behavior. Moral Judgments as Forms of Self-guidance In Kohlberg s model, which assumes that all the moral judgments people make stem from the same structure of moral judgment, there is no reason to expect any differences between the moral judgments people make to and about themselves and the moral judgments they make to and about others. In contrast, from an evolutionary perspective, we would expect the form of the two types of judgment to differ in significant ways. We would expect the judgments people make about their rights and duties to be more selfish than the judgments they make about the rights and duties of others for two reasons. First, we would expect people to be motivated to advance their own interests. Second, we would expect the authors of self-serving moral judgments to constitute more receptive audiences than most other recipients. Thus, we would expect people to send messages to themselves such as, you deserve more than your share, your contributions are worth more than those of others, and you should look after your own interests, then graciously receive them with favor. The magnitude of self-serving biases in moral judgment should, however, vary across a variety of conditions. First, when people express moral judgments about themselves to others, recipients should exert a constraining effect on the selfishness of the judgments in much the same way they exert a constraining effect on the selfishness of second-person moral judgments. We would expect audiences to call people on their selfishness and point out inconsistencies in the standards they are applying to themselves and to others. For this reason, in general, we would expect people to make less selfish moral judgments in public than in private. People also should

18 Evolutionary reconceptualization of Kohlberg 18 be selective in the audiences to whom they express their moral judgments, choosing those they anticipate will be most receptive to their points of view (Krebs & Denton, 1997). Second, the extent to which people apply to themselves the standards they preach to others may depend on the persuasiveness of their communications. When individuals adduce arguments in support of the moral judgments they make to others, they may in effect persuade themselves that the arguments are right and buy into them (see Brown, 1986; Taylor, 1989). As suggested by Trivers (2000), believing in the standards one preaches to others may enhance the persuasive power of one s communications. Alternatively, people may compartmentalize moral judgments about themselves and others. Third, the mechanisms through which people make moral judgments may be designed in ways that contain cognitive representations of others. In effect, internalized images of others may send and receive moral messages. Freud has suggested that introjected images of parents, or more exactly their superegos, tell people what they ought to do and sit in judgment of them when they fail to live up to their standards. The idea that the psychological mechanisms through which people make moral judgments to themselves, often called their conscience, contain internalized representations of others, especially their parents, has been advanced by scholars from a wide variety of theoretical traditions (e.g., Aronfreed, 1968; Higgins & Eccles-Parson, 1983). However, in the same way that people may direct their moral judgments to audiences who are receptive to their biases, they may invoke internalized audiences who exhort them to advance their own interests and help them justify their self-serving behaviors. In the end, we must rely on empirical research to map the design of structures that give rise to moral judgments about selves and others. Although a host of studies have reported selfserving biases in social and moral judgments (see Krebs, Denton & Higgins, 1988; Krebs &

19 Evolutionary reconceptualization of Kohlberg 19 Denton, 1997; MacDonald, 1988; Taylor, 1989, for reviews), people differ in the extent to which they harbor such biases, and the biases tend to diminish, disappear, and even reverse themselves when people make judgments about their friends and relatives (Krebs & Laird, 1998). The Evolution of Moral Behavioral Dispositions Implicit in the analysis of the evolution of moral judgment I have been advancing is a profoundly important implication, namely that humans are biologically endowed with the capacity to behave in moral ways. If, as I have argued, people use moral judgments such as those classified by Kohlberg and his colleagues to activate evolved mechanisms that induce recipients to behave in accordance with their prescriptions, recipients must possess such evolved mechanisms. Indeed, I would expect the behavior-producing mechanisms to have evolved before the judgment-producing mechanisms, else the latter would have nothing to activate. But could mechanisms that dispose individuals to behave morally have evolved? conclusions: Some very eminent evolutionary theorists think not. Consider for example the following Nothing resembling the Golden Rule or other widely preached ethical principles seems to be operating in living nature. It could scarcely be otherwise, when evolution is guided by a force that maximizes genetic selfishness (Williams, 1989, p. 195) Be warned that if you wish, as I do, to build a society in which individuals cooperate generously and unselfishly toward a common good, you can expect little help from biological nature. Let us try to teach generosity and altruism, because we are born selfish. (Dawkins, 1976, p. 3) As much as I admire the work of these theorists, I believe their conclusions about the evolution of morality are incorrect, or at least misleading. When evolutionary biologists talk about selfish behaviors, they mean behaviors that help individuals propagate their genes. Such theorists are, in effect, saying that the mechanisms that give rise to all evolved behaviors (that is

20 Evolutionary reconceptualization of Kohlberg 20 to say, behaviors that helped ancestors propagate their genes) are designed in ways that help individuals propagate their genes in modern environments. But this is not necessarily true, because, among other reasons, behaviors that were adaptive, or genetically selfish, in ancestral environments may not be adaptive in current environments. Further, genetically selfish behaviors are not necessarily individually selfish, which is the criterion most people use for morality. As examples, it may be genetically selfish to sacrifice oneself for the sake of one s offspring or other kin, but such behaviors would not be considered individually selfish or immoral. Although cooperative behaviors may foster the genetic interests of cooperators (rendering cooperation genetically selfish), such behaviors may nevertheless qualify as moral. In my view, there is no necessary inconsistency between behaving morally and fostering one s biological interests. It is not immoral to attempt to survive or to reproduce; indeed some think people have a moral obligation to preserve their lives and to bear children. The criterion for morality pertains to how we go about fostering our biological interests. In general, we consider it moral to foster our biological interests in ways that foster the interests of others that is to say in cooperative ways and we consider it immoral to foster our interests at the expensive of others that is to say in selfish ways. I believe there is little question that mechanisms giving rise to the cooperative strategies prescribed by Kohlbergian Stage 1, 2 and 3 moral judgments have evolved in the human species, and in some other species. I also believe that such mechanisms can, under certain conditions, give rise to Stage 4 and Stage 5 moral strategies. I have reviewed evidence supporting this claim in several publications (Krebs, 1998; Krebs, 2000a, b, c; Krebs & Janicki, 2003). I will briefly summarize it here. The Evolution of Stage 1 Strategies Stage 1 moral judgments prescribe obeying those who are older, bigger, or more

21 Evolutionary reconceptualization of Kohlberg 21 powerful than you and the rules they enforce in order to avoid getting beaten up, hit, punished, put in jail, or killed. It is easy to see how the strategy implicit in such judgments could pay off better than more selfish attempts to maximize one s gains. Members of groups who are relatively weak face a Hobson s choice: either defer to those who are more powerful, or suffer the consequences. Faced with such choices, it is more adaptive for relatively subordinate members of groups to submit to the authority of the more dominant members in order to make the best of a bad situation and live to fight another day. There is a spate of evidence that mental mechanisms giving rise to submissive and subordinate behaviors have evolved in many species (see Alcock, 1998; Boehm, 2000; Krebs, 1998, 2000b; Sloman & Gilbert, 2000), including humans (Milgram, 1964; Strayer & Strayer,1976). In some species, such mechanisms produce dominance hierarchies or pecking orders. Deferential mechanisms are rooted in fear systems, combined perhaps with a touch of awe. The Evolution of Stage 2 Strategies Stage 2 moral judgments prescribe helping other people, because you may need them to do something for you one day, keeping promises so the other person will keep promises to you or give you something in return, and so on. In short, you scratch my back and I ll scratch your back. As pointed out by Trivers (1971) and others, individuals who reciprocate resources may gain more than individuals who do not through gains in trade. In some contexts individuals are able to bestow huge benefits on others at very little cost to themselves. Trivers (1971) gives the example of someone throwing a drowning person a life saver. To evolve, Stage 2 cooperative strategies must contain antidotes to exploitation. One of the strategies prescribed by Stage 2 moral judgments is commonly called Tit for Tat. Tit for Tat

22 Evolutionary reconceptualization of Kohlberg 22 is based in the decision-rule, make a cooperative overture, then copy the response of your partner in subsequent exchanges. If your partner cooperates, you glean the benefits of the cooperative exchange. If your partner behaves selfishly, you cut your losses (and his or hers!) by behaving selfishly until he or she relents and behaves cooperatively. This aspect of the strategy is prescribed by Stage 2 moral judgments such as an eye for an eye and don t get mad, get even. Computer simulations of the evolution of social strategies have found that Tit for Tat can defeat more selfish strategies if it enters populations in sufficiently large numbers (Axelrod & Hamilton, 1981). Although Tit for Tat always loses to selfish strategies one on one, it ends up winning evolutionary contests by cutting its losses in exchanges with selfish strategies and reaping the benefits of drawing with cooperative strategies, including itself. (When strategies evolve, they become increasingly prevalent in populations; thus, evolved social strategies are usually designed in ways that enable them to do well in exchanges with replicas of themselves.). Trivers (1971), Dugatkin (1997) and others have adduced evidence demonstrating that mechanisms giving rise to reciprocity have evolved in several species. The exchange of blood in vampire bats is a particularly interesting case in point (Wilkinson, 1990). Gouldner (1960) and others have suggested that reciprocity is a universal moral norm in the human species. As suggested by Trivers (1971) reciprocity is rooted in human psychological systems giving rise to a sense of deserving, a sense of gratitude and indebtedness, and senses of righteous indignation, retribution, revenge, and vindictiveness as well as systems instilling a sense of fairness and justice. The Evolution of Stage 2/3 Strategies Stage 2/3 strategies prescribe giving those who have transgressed against you a second chance if they have suffered enough or if they feel bad or sorry about cheating you. Such

23 Evolutionary reconceptualization of Kohlberg 23 strategies are reflected in aphorisms such as forgive and forget and everyone makes mistakes. Following the publication of Axelrod and Hamilton s (1981) findings, investigators changed parameters in Axelrod and Hamilton s computer simulations and examined the fecundity of other strategies. In general, the closer the environments approximated the actual conditions of evolution, the greater were the benefits from conditionally altruistic strategies. For example, games that allowed for the inevitable errors that occur in social exchanges found that strategies such as Two Tits for a Tat, Generous Tit for Tat, Contrite Tit for Tat, and Forgiving Tit for Tat fared better than Tit for Tat because they were equipped to break the selfdefeating blood feuds precipitated by one selfish mistake (see Krebs, 2000; Ridley, 1996, for reviews of relevant research). Simulations that enabled players to observe other players and keep track of their strategies (called image scoring ) favored the evolution of altruism through indirect reciprocity (Nowak & Sigmund, 1998). The Evolution of Stage 3 Strategies Among the Stage 3 moral judgments classified in Kohlberg s system, there are at least two distinguishable types of strategy. Translated into biological terms, the first one prescribes enhancing one s inclusive fitness by helping members of one s group. The second prescribes conforming to moral norms. The adaptive benefits of the first type of strategy vary in accordance with the relationship between the helper and the recipient of help. Helping relatives. Many of the moral judgments contained in Kohlberg s scoring manual prescribe upholding families and helping relatives. It is easy to account for the evolution of mechanisms giving rise to such strategies. It is quite common among mammals for parents to sacrifice their somatic interests for the sake of their offspring. Such self-sacrificial behaviors

24 Evolutionary reconceptualization of Kohlberg 24 help the parents propagate their genes. In a classic paper, Hamilton (1964) pointed out that the biological value of parental investment can be extended to relatives other than offspring. The probability of individuals sharing genes varies in proportion to their degree of relatedness. For example, the probability of parents and their offspring sharing genes is.5. Same for siblings. The probability of grandparents and their grandchildren sharing genes is.25; and so on. Hamilton (1964) explained how a decision-rule could evolve that induced individuals to help others when the coefficient of their relatedness is greater than the cost to the helper of helping, divided by the benefits to the recipient (r>c/b). Alcock (1998) and others have reviewed evidence that members of many species behave in accordance with the strategy implicit in Hamilton s equation. Helping individuals who seem similar to relatives. Mechanisms that direct individuals to help their relatives are limited morally because they give rise to nepotistic and discriminatory behaviors, inducing individuals to favor relatives over non-relatives, close relatives over distant relatives, and more fecund relatives over less fecund relatives. This helps explain the imbalance of helping between parents and children, and grandparents and grandchildren. However, perhaps fortunately, animals may not be able to detect the proportion of genes they share with others directly or exactly (Dawkins, 1989, but see Pfennig & Sherman, 1995 and Rushton, 1999). Animals may have to rely on kin recognition cues such as phenotypic similarity, familiarity, and proximity (Porter, 1987). Although such cues may have been highly predictive of genetic relatedness in ancestral environments, they may activate mechanisms that induce individuals to help non-relatives in modern environments. Inasmuch as the genetic payoffs to individuals from helping non-relatives are less than those from helping their relatives, helping non-relatives can be considered more altruistic than helping relatives (see Krebs, 1987).

25 Evolutionary reconceptualization of Kohlberg 25 Helping friends. Many Kohlbergian moral judgments prescribe helping friends. In part, the evolution of mechanisms that dispose individuals to help their friends can be accounted for in terms of the misfiring of kin-selected strategies. Friends usually resemble kin; friends tend to be similar to one another, familiar, and live in relatively close proximity. In addition, such mechanisms probably reaped adaptive benefits through more complex forms of reciprocity than those prescribed by Stage 2 moral judgments. Tooby and Cosmides (1996) have argued that exchanges between friends do not usually conform to decision-rules such as those that define Tit for Tat strategies. Friends do not pay each other back for every favor they bestow on one another, and they do not typically seek revenge when their partners fail to repay every debt. The give and take between friends is more general, averaged over many kinds of resource and over long periods of time. Tooby and Cosmides (1996) discuss the biological significance of a phenomenon they call the banker s paradox : banks are least likely to lend people money when they most need it and are least likely to be able to pay it back. In Tooby and Cosmides view, investing in friends is a strategy aimed at increasing the probability that people will have bankers who are willing to lend them money when they are in need. Accumulating resources counts for nothing in evolution if they do not enhance individuals chances of surviving, reproducing, and propagating their genes. Bestowing relatively low cost favors on friends over long periods of time can be a winning strategy if it induces them to help save your life, help you find a mate, or foster the fitness of your relatives. In this sense, helping friends is akin to investing in insurance. Helping In-group members. Kohlbergian moral judgments also prescribe helping members of one s in-group. Such strategies could stem from mechanisms of social control, which I discuss later, from extensions of kin-selected mechanisms and from extensions of

26 Evolutionary reconceptualization of Kohlberg 26 mechanisms designed to reap benefits from reciprocity. With respect to extensions of kinselected mechanisms, Sober and Wilson (1998) explain how, under certain (admittedly rare) circumstances, mechanisms disposing individuals to behave in genetically altruistic ways that is to say, in ways that enhance the fitness of members of their group at the expense of their own inclusive fitness can evolve when the group as a whole benefits more than groups containing less altruistic members. The evolutionary process involved is similar to kin selection inasmuch as it is based in propagating genes that dispose individuals to behave altruistically by helping other individuals who possess replicas of such genes. However, it differs from kin selection, because the mechanisms that mediate altruism are not calibrated in ways that favor those who are genetically-similar to the altruist. In group selection, the costs of helping those who do not possess the genes that give rise to altruism are outweighed by the benefits to the group as a whole, and selfish members are reduced in proportion when groups recombine (see Sober & Wilson, 1998). Mechanisms that induce individuals to help members of their groups also could have evolved through the benefits of complex systems of reciprocity. Alexander (1987) has argued that the biological function of moral dispositions is to uphold systems of indirect reciprocity. Such systems may assume two forms. In the first, person A helps person B; person B helps person C; and person C helps person A. In the second, all members of a group contribute resources to a central distributor who redistributes the resources to those who have contributed. Systems of indirect reciprocity have significantly more potential than systems of direct exchange to maximize benefits for all participating members, because they enable each individual to obtain resources of maximum value to them and minimum cost to those who produced them in exchange for resources of minimum cost to them and maximum benefit to recipients. Such

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