The Nervous System. Autonomic Division. C h a p t e r. PowerPoint Lecture Slides prepared by Jason LaPres North Harris College Houston, Texas

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C h a p t e r 17 The Nervous System Autonomic Division PowerPoint Lecture Slides prepared by Jason LaPres North Harris College Houston, Texas Copyright 2009 Pearson Education, Inc., publishing as Pearson Benjamin Cummings

Introduction Routine adjustments in physiological systems are made by the autonomic nervous system (ANS). The ANS regulates body temperature and coordinates cardiovascular, respiratory, digestive, excretory, and reproductive functions. It adjusts internal water, electrolyte, nutrient, and dissolved-gas concentrations in body fluids outside our conscious awareness.

A Comparison of the Somatic and Autonomic Nervous Systems The autonomic nervous system differs from the somatic nervous system in the arrangement of the neurons connecting the central nervous system to the effector organs. Visceral motor neurons in the CNS, known as preganglionic neurons, send their axons, called preganglionic fibers, to synapse on ganglionic neurons, whose cell bodies are located outside the CNS, in autonomic ganglia. Axons from the ganglionic neurons are called postganglionic fibers because they carry impulses away from the ganglion. Postganglionic fibers innervate peripheral tissues and organs, such as cardiac and smooth muscle, adipose tissue, and glands.

A Comparison of the Somatic and Autonomic Nervous Systems Subdivision of the ANS Sympathetic division Generally predominant under resting conditions Parasympathetic division Generally kicks in during times of exertion, stress, or emergency Both divisions affect their target organs through the release of neurotransmitters by postganglionic fibers

A Comparison of the Somatic and Autonomic Nervous Systems Figure 17.1a Components and Anatomic Subdivisions of the ANS (Functional Components)

A Comparison of the Somatic and Autonomic Nervous Systems Figure 17.1b Components and Anatomic Subdivisions of the ANS (Anatomical Subdivisions)

The Sympathetic Division The sympathetic division consists of the following: Preganglionic neurons located between segments of the spinal cord Two types of ganglionic neurons in ganglia near the vertebral column Sympathetic chain ganglia, also called paravertebral, or lateral ganglia Collateral ganglia, also known as prevertebral ganglia Specialized neurons in the interior of the suprarenal gland

The Sympathetic Division Figure 17.2 Organization of the Sympathetic Division of the ANS

The Sympathetic Division Figure 17.3a Sympathetic Pathways and Their General Functions (Sympathetic Chain Ganglia)

The Sympathetic Division Figure 17.3b Sympathetic Pathways and Their General Functions (Collateral Ganglia)

The Sympathetic Division Figure 17.3c Sympathetic Pathways and Their General Functions (The Suprarenal Medullae)

The Sympathetic Division Figure 17.4 Anatomical Distribution of Sympathetic Postganglionic Fibers

The Sympathetic Division Figure 17.5 Suprarenal Medulla

The Sympathetic Division Figure 17.6 Sympathetic Postganglionic Nerve Endings

The Sympathetic Division In summary: The sympathetic division of the ANS includes two sympathetic chains, three collateral ganglia, and two suprarenal medullae. Preganglionic fibers are short because the ganglia are close to the spinal cord. The sympathetic division shows extensive divergence. All preganglionic neurons release ACh at their synapses with ganglionic neurons. The effector response depends on the function of the membrane receptor

The Sympathetic Division Innervation Patterns

The Parasympathetic Division The parasympathetic division of the ANS includes the following: Preganglionic neurons located in the brain stem and in sacral segments of the spinal cord Ganglionic neurons in peripheral ganglia located very close to or even within the target zones As a result, effects are more specific and localized than those of sympathetic division

The Parasympathetic Division Figure 17.7 Organization of the Parasympathetic Division of the ANS

The Parasympathetic Division Figure 17.8 Anatomical Distribution of the Parasympathetic Output

The Parasympathetic Division General functions of the parasympathetic division: Constriction of the pupils to restrict the amount of light entering the eyes; assists in focusing on nearby objects Secretion by digestive glands, including salivary glands, gastric glands, duodenal and other intestinal glands, the pancreas, and the liver Secretion of hormones that promote nutrient absorption by peripheral cells Increased smooth muscle activity along the digestive tract Stimulation and coordination of defecation

The Parasympathetic Division General functions of the parasympathetic division (continued) Contraction of the urinary bladder during urination Constriction of the respiratory passageways Reduction in heart rate and force of contraction Sexual arousal and stimulation of sexual glands in both sexes

The Parasympathetic Division In summary: The parasympathetic division includes visceral motor nuclei in the brain stem associated with four cranial nerves (III, VII, IX, and X). The ganglionic neurons are situated in intramural ganglia or in ganglia closely associated with their target organs. The parasympathetic division innervates structures in the head and organs in the thoracic and abdominopelvic cavities. All parasympathetic neurons are cholinergic. The effects of parasympathetic stimulation are usually brief and restricted to specific organs and sites.

The Parasympathetic Division Innervation Patterns

Relationships between the Sympathetic and Parasympathetic Divisions The divisions of the autonomic nervous system are not isolated. Sympathetic and parasympathetic divisions work together: Dual innervation Peripheral autonomic plexuses

MRelationships between the Sympathetic and Parasympathetic Divisions Figure 17.9 The Peripheral Autonomic Plexuses

MRelationships between the Sympathetic and Parasympathetic Divisions Figure 17.10 A Comparison of the Sympathetic and Parasympathetic Divisions

MRelationships between the Sympathetic and Parasympathetic Divisions TABLE 17.1 A Comparison of the Sympathetic and Parasympathetic Divisions of the ANS

Integration and Control of Autonomic Functions The ANS is organized into a series of interacting levels Visceral reflexes Short reflexes Long reflexes Enteric nervous system (ENS) Higher levels of autonomic control Hypothalamus

Integration and Control of Autonomic Functions Figure 17.11 Visceral Reflexes

Integration and Control of Autonomic Functions TABLE 17.2 Representative Visceral Reflexes

Integration and Control of Autonomic Functions Figure 17.12 Levels of Autonomic Control