(For the Students of B.V.Sc. & A.H. Degree Programme)

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m m m j ilt L Ijy! ]H 3 Q fflq Q Q a i3 (D II ] (For the Students of B.V.Sc. & A.H. Degree Programme)

(For the Students of B.V.Sc. & A.H. Degree Programme) M.S.Saxena CBS PUBLISHERS & DISTRIBUTORS PVT. LTD. NEW DELHI BENGALURU. CHENNAI. KOCHI. MUMBAI. PUNF

ISBN : 81-239-0716-8 First Edition: 2000 Reprint: 2004, 2007, 2009, 2011, 2012, 2015 Copyright by the Author & Publisher All rights reserved. No part o f this book may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopying, recording, or any information storage and retrieval system without permission, in writing, from the authors. Published by: Satish Kumar Jain for CBS Publishers & Distributors Pvt. Ltd., CBS Plaza, 4819/XT Prahlad Street, 24 Ansari Road, Daryaganj, New Delhi - 110002, India delhi@ cbspd.com, cbspubs@ airtelmail.in www.cbspd.com Ph.: 23289259, 23266861, 23266867 Fax: 0! 1-23243014 Branches. Bengaluru: Seema House, 2975, 17th Cross, K.R. Road, Bansankari 2nd Stage, Bengaluru - 560070 Ph: +91-80-26771678/79 Fax:+91-80-26771680 E-mail: cbsbng@gmail.com, bangalore@cbspd.com Chennai: No. 7, Subbaraya Street, Shcnoy Nagar, Chennai - 600030 Ph: +91-44-26681266,26680620 Fax:+91-44-42032115 E-mail: chennai@cbspd.com Kochi: 36/14, Kalluvilakam, Lissie Hospital Road, Kochi - 682018 P h :+91-484-4059061-65 Fax:+91-484-4059065 E-mail: cochin@cbspd.com Mumbai: 83-C, Dr, E. Moses Road, Worli, Mumbai - 400018 P h :+91-9833017933,022-24902340/41 E-mail: mumbai@cbspd.com Pune: Bhuruk Prestige, Sr. No. 52/12/2+1+3/2, Narhe, Haveli (Near Katraj-Dehu Road Bypass), Pune - 411041 P h :+91-20-64704058/59,32342277 E-mail: pune@cbspd.com Printed a l : J.S. Offset Printers, Delhi

Foreword r There is a great paucity of textbooks in the field of veterinary sciences written by Indian authors as reference and textbooks suitable to Indian college curriculum. Most Indian teachers use textbooks written by American and European authors directed at the clientele in their respective countries wh.ch are supplemented by notes to suit experience of Indian and other developing countries. This is a very unsatisfactory situation. In spite of a number of efforts by Indian Council o! gricultural Research and the publishing community at large, the number of authors who have u*ken up the challenge of writing books to serve as textbooks in veterinary colleges are very few. There seems to be no tradition which could generate a crop of authors who could take upon themselves the very important task of writing textbooks suitable for our curriculum in the veterinary colleges. With the notification of standards for veterinary education by the Indian Veterinary Council, the entire course curriculum has been revised/updated and it is therefore necessary that attempts sho'ild be made by various university teachers to prepare texts to suit this course curriculum. Dr. Saxena s attempt to produce a basic text on andrology and artificial insemination based on the syllabus prescribed by Indian Veterinary Council is a good attempt in this direction. The need for a comprehensive book on clinical andrology and artificial insemination meant for undergraduates in the veterinary schools was urgent; this book can now be used as a textbook which will fill the long felt need in the subject. The text is primarily devoted to andrological examination of breeding bulls and their management, collection, evaluation and preservation of semen particularly of buffaloes. This is going to be very useful to the beginners. A chapter on cleaning and sterilization of equipment, recording of data is important from the point of view of veterinary practice. I believe tha, this compendium will be used as a workbook by the teachers as well as students. They should assist Dr. Saxena in improving its functional use in due course of time. P.N. Bhat Chairman, World Buffalo Trust

Preface Reproduction is a joint venture in which both males and females participate. Reproductive disorders are more apparent in females, hence failures are often attributed to female gender only. A close examination of the process, however, reveals that males are as great offenders as females to cause infertility problems. The male should provide sperms which are competent to fertilize the ovum. Whenever there is large-scale problem of infertility in a herd or in large number of females, the male is always seen with suspicion. In addition to productive efficiency, the clinical examination of the bull for its reproductive efficiency is gaining more and more popularity. There are many excellent books on andrology and A.I. which would undoub <ly remain unmatched for many years to come. None of these are tailor-made for the B.V.Sc. & A H. syllabus prescribed by Indian Veterinary Council and followed by Indian veterinary institutions. Necessity for a concise book on Clinical Andrology and A.I. within the scope and apprehension of undergraduate veterinary students has thus been felt for long. This compendium cum workbook has been attempted with the expectation that it would fulfill this long felt need and would provide necessary guidelines to the B.V.Sc. & A.H. students. Basically, the contents of the book have been drawn from diverse literary sources as well as the practical field experience of the author. There might be some errors in the text and there is tremendous scope for improvement. The suggestions and criticisms would always be heartily welcomed. Many of my senior colleagues, my associates, friends and students have made valuable contributions towards compilation of this book and the author is deeply indebted to Dr. Harpal Singh, Dean, College of Veterinary Sciences; Dr. S.N. Maurya, Prof. & Head, Department of Gynaecology and Obstetrics; Dr. B.D. Lakhchaura, Prof. & Head, Department of Veterinary Biochemistry for their insj iralion and valuable suggestions. Dr. V. Umapathi, Junior Research Officer and Mr. K.C. Joshi, Steno-typist of the Department of Veterinary Biochemistry deserve exceptional appreciation for their wholehearted help. Sincere thanks are due to Dr. V.B. Saxena, Dr. H.P. Gupta and Dr. Shiv Prasad for their critical suggestions. Km. Geeta Kandpal, Teaching Associate of the Department of Veterinary Biochemistry deserves appreciation in handling work on computer. The author would like to express his appreciation and indebtedness to his wife Meena and daughters, Shilpi and Shubhangi for their understanding and forgiveness they had shown for his several long absences from home. M.S. Saxena vii

Syllabus (For B.V.Sc. & A.H. Degree Programme) Andrology and Artificial Insemination VO G -511 Cr. H rs. 2 + 0 = 2 Introduction, development, comparative study of male genitalia and gonads. Growth, puberty, sexual maturity and libido. Endocrine control of reproduction in the male domestic animals. Factors affecting maturity and sex drive in bulls. Sexual behaviour in males. Forms of male infertility. General considerations. Factors affecting infertility in male, its treatment and diagnosis. Diseases, abnormalities and malformations of male genitalia, their diagnosis and treatment of coital injury and infections. Testicular hypoplasia and degeneration. Diseases of the accessory sex glands. Introduction, history, development, advantages and limitations of A.I. methods of semen collection in various species. Technique of A.I. factors affecting quality and quantity of semen. Tests for evaluation of semen; extension of semen; preservation of semen at different temperatures. Storage and shipment of semen, metabolism of semen, biochemistry of semen. Andrology and Artificial Insemination (Clinics) VO G -512 C r. H rs. 0 + 2 =: 2 Andrological investigations of breeding bulls. Assessment of sires. Physical examinations observing sexual behaviour; palpation of scrotum, spermatic cord, seminal vesicles and ampullae. Collection of materials for sperm activity, morphology and diagnosis of reproductive disorders in bulls. Preparation of A.V., collection of semen, evaluation, dilution, preservation techniques at different temperature. Freezing of semen. Insemination techniques in chilled and frozen semen. Planning and organization of A.I. centre. Selection, care, training and maintenance of breeding bulls for A.I., recording systems. Care, sterilization. Storage and upkeep of equipments used for Artificial Insemination. viii

Contents Foreword Preface. Syllabus. VII v VllI SECTION I 1. Reproductive organs of m ale dom estic animals 1 Scrotum and testes 1 Thermoregulation of testes 6 Functions of the testes 7 Epididymis 7 Functions of the epididymis 7 Vas deferens 8 Accessory sex glands 9 The vesicular glands (seminal vesicles) 9 The prostate gland 9 The bulbourethral glands (Cowper s glands) 10 Urethra 10 Penis 10 Prepuce 13 Blood and nerve supply 13 2. The em bryology of the m ale reproductive tract... 14 Gonadogenesis in males 15 The indifferent stage 15 Testes 15 Ovary 17 ix

x Contents Descent of the testicles 17 The origin of the internal ducts 19 Origin of the accessory sex glands 21 Origin of the external genitalia 21 3. G row th, puberty, sexual m aturity and l i b i d o...23 Hormonal changes 24 Spermatogenesis 24 Anatomical changes 24 Testicular development 24 Factors affecting age at puberty and sexual maturity 24 Genetic factors 25 Nutritional factors 25 Hormonal factors 25 Social and climatic factors 25 Body weight 25 Libido 25 Hereditary factors 26 Nutrition 26 Systemic diseases 26 Age 26 Management 27 Psychogenic factors 27 Climatic factors 27 Endocrine factors 27 Injuries of joints, muscles, nerves and tendons 27 Diseases of the penis and prepuce 27 4. Endocrine control of reproduction in male dom estic a n im a ls...28 Testicular descent 28 Before puberty 28 GnRH and gonadotropins 28 The Leydig s cells and Sertoli cells 29 The steroid hormones 30 Steroidogenesis 30 Physiological actions of androgens 31 Other hormones 34 5. Sexual behaviour in male a n i m a l s...35 Sexual arousal 37 Courtship (sexual display) 37 Erection and penile protrusion 37

Contents Mounting 37 Intromission 38 Ejaculation 38 Dismounting 39 Refractoriness 39 Infertility in male a n im a ls...40 Forms of infertility in males 40 Impotentia coeundi 40 Baianoposthitis 43 Phimosis 44 Paraphimosis 44 Diphallus 44 Phallocampsis 44 Adhesions of the penis and prepuce 45 Ruptured or broken penis 45 Tumors of the penis and prepuce 45 Chronic prolapse of the prepuce 46 Other causes of the loss of libido and inability to copulate 46 Hernias 46 Premature erection 46 Loss of sensory innervations of the glans penis 46 Urinary caliculi 46 Pain yielding causes 46 Impotentia generandi 47 Impotentia generandi associated with apparently normal semen 47 Infectious diseases 47 Inherited sperm defects 47 Diadem effect 47 Knobbed spermatozoa 47 Gene or chromosomal defects 47 Atypical basic nuclear proteins 47 Enzymatic disturbances 47 Impotentia generandi assiciated with apparently abnormal sperm 48 Cryptorchidism 48 Imperfect descent of testes 48 Scrotal or inguinal hemia 48 Testicular hypoplasia 48 Testicular degeneration 49 Orchitis 53 xf

Veterinary Andrology & Artificial Insemination Publisher : CBS Publications ISBN : 9788123907161 Author : Saxena MS Type the URL : http://www.kopykitab.com/product/7748 Get this ebook