RESEARCH ARTICLE. Nineteen new species of Taperinha (Hemiptera: Cicadellidae: Deltocephalinae)

Transcription

1 RESEARCH ARTICLE TAES 141: ISSO Nineteen new species of Taperinha (Hemiptera: Cicadellidae: Deltocephalinae) Paul H. Freytag Emeritus Professor, Department of Entomology,University of Kentucky, Lexington, Kentucky Abstract Nineteen new species are added to the Genus Taperinha Linnavouri from South America. The new species are Taperinha nuda, T. trigona, T. angulata, T. hamata, T. media, T. ungulata, T. lobata, T. anisota, T. lapayensis, T. odonta from Colombia; T. retusa, T. zanolae from French Guiana; T. dozieri, T. longicornis from Ecuador; T. elongata from Suriname; T. stypa from Venezuela; and T. bulba, T. stenostyla, T. peruviana from Peru. A key to the species known from Colombia is included. Key Words: Hemiptera, Cicadellidae, Deltocephalinae, Taperinha, South America, new species. INTRODUCTION The genus Taperinha was described by Linnavuori (1959) and included 3 new species. Linnavuori & DeLong (1978) added 3 more new species. Zanol (1988, 1989, 1997, and 2004) added 17 new species. In this paper 19 new species are described, all species in the genus are from South America, and only one species (T. punctata Linnavouri) is also found in the southern part of Central America (Costa Rica & Panama). With the new species there are now 42 species known in this genus. A key to the species found in Colombia is included after the description of the new species. The species of Taperinha can be identified by the following external characters: The crown of the head is usually light yellow with two large dark spots, with a distinct anterior margin. The spots are usually black, but can also be brown, red or orange. The forewings have extra cross veins on the costal margin, and with or without a saddle-like pattern across the claval areas. The underside including the face is dark brown or black. They are usually from 4 to 5 mm. in length, and somewhat oval in dorsal view. These characters are somewhat distinct and unusual for species in the Deltocephalinae in South America. The acronyms used in this paper are the following: IHVL = Instituto von Humboldt, Villa de Leyva, Colombia UKYL = University of Kentucky Collection, Lexington, Kentucky, USA MNHN = Musėum national d Histoire naturelle, Paris, France FSCA = Florida State Collection of Arthropods, Gainesville, Florida, USA MUSM = Museo de Historia Natural, Lima, Peru. A list of the species of this genus and the known distribution, and a key to the species known to occur in Colombia are given after the descriptions of the new species. Taperinha nuda Freytag, new species (Figures 1-5) urn:lsid:zoobank.org:act:2324bdf2-d227-4e6d-92c3-dcbf6f99ac87 Description Length of males mm., females mm. Head with crown yellow, with four evenly spaced small black spots across anterior margin, pair of small light brown spots behind median pair of black spots, posterior half with two large brown spots. Pronotum yellow across anterior Publication date: 31 December 2015 urn:lsid:zoobank.org:pub:dbbdac30-ceb4-4ee d9ecd9d565fb

2 464 TWENTY-ONE NEW SPECIES OF TAPERINHA margin, with some brown spots, posterior two thirds dark brown. Scutellum yellow, with two small dark brown spots on each lateral margin, pair of similar spots in middle, with triangles light brown. Forewings light brown, with distinct brown saddle, costal margin with many dark brown cross veins. Ventral surface dark brown. Male genitalia: Pygofer (Fig.1) broadly rounded at apex, without processes. Subgenital plate (Figs. 1 & 2) longer than pygofer, triangular in ventral view. Style (Fig. 2) short, broad at base, hooked at apex, less than half length of subgenital plate. Connective (Figs. 3 & 4) nearly same length as aedeagus. Aedeagus (Figs. 3 & 4) narrow, without processes, gradually narrowing to pointed apex. Female genitalia: Seventh sternum (Fig. 5) nearly rectangular, but with slightly narrower, evenly rounded posterior margin, with peg-like extension on each side. Type data Holotype male: Colombia: Valle del Cauca, PNN Farallones de Cali, 3 26 N W, 650 m., Malaise, 30-I-13-II-2001, S. Sarria Leg., M1528 (IHVL). Paratypes: Three males, one female, same data as holotype, except III-2001, M1527 (one male, one female- IHVL, two males-ukyl). Other specimens seen One male, same data as holotype, except 9-V-18-VII-2000, M1100 (IHVL); two males, same data as holotype, except 27-II-27-III-2001, M1536 (IHVL); one male, same data as holotype, except 21-XI-19-XII-2000, M1541 (IHVL); one male, same data as holotype, except 2-16-I-2001, M1544 (IHVL); one male, same data as holotype, except I-2001, M1547 (IHVL); one male, same data as holotype, except 27-III-24-IV-2001, M1899 (IHVL); one male, same data as holotype, except VII-2001, M2837 (IHVL); one male, same data as holotype, except 24-IV-8-V-2001, M1900 (IHVL); one female, same data as holotype, except 900 m., 27-II-27-III-2001, M1532 (IHVL); one female, same data as holotype, except 730 m., 19-VI-3-VII-2001, M1889 (IHVL); one female, same data as holotype, except 900 m., 19-VI-3-VII-2001, M1891 (IHVL); one female, same data as holotype, except 8-V-19-VI-2001, M1892 (IHVL); two females, same data as holotype, except 730 m., 8-V-19-VI-2001, M1893 (one-ihvl, one UKYL); one female, same data as holotype, except 900 m., 8-V-19-VI-2001, M1894 (IHVL); two females, same data as holotype, except Anchicaya, 730 m., 3-17-VII-2001, M2833 (one-ihvl, one-ukyl); one male, same data as holotype, except Anchicaya, VII-2001, M2837 (IHVL); one male, same data as holotype, except Anchicaya, 730 m., 28-VIII-11-IX-2001, M2864 (IHVL); one female, same data as holotype, except Anchicaya, 730 m., 11-IX-16-X-2001, M2867 (IHVL); one male, same data as holotype, except Anchicaya, 900 m., M2868 (IHVL); one male, Colombia: Nariño, RN La Plenada, Percela Olga, 1 15 N 78 15W, 1850 m., Malaise, 2-16-VI- 2000, G. Oliva Leg., M250 (IHVL); one male, same data, except Via Hondón, 1930 m., 2-16-IX-2000, M885 (IHVL); one male, same data, except Via Hondón, 16-VIII-2-IX-2000, M917 (IHVL); and one male, same data, except Parcala Olga, 1850 m., , M2363 (IHVL). Notes This species is similar to punctata in size and color pattern. It differs in both the male and female genitalia. In nuda the male pygofer lacks a ventral process, and in punctata the male pygofer has one process. The female seventh sternum is convex medially on the posterior margin for nuda, and slightly concave for punctata. Etymology This species is named for the lack of a male pygofer process. Taperinha trigona Freytag, new species (Figures 6-10) urn:lsid:zoobank.org:act:cece72e6-ecb2-4f2d-8a84-b3a d4 Description Length of males mm., females mm. Head with crown yellow, with four faint small black spots across anterior margin, posterior half with two large light brown spots. Pronotum and scutellum light brown with numerous dark brown spots. Forewings light brown, with very small brown speckles overall, costal margin with numerous black cross veins. Ventral surface dark brown. Male genitalia: Pygofer (Fig. 6) truncate at apex, with triangular process from posterior ventral margin extending dorsally to anal tube. Subgenital plate (Figs. 6 & 7) longer than pygofer, triangular in ventral view. Style (Fig. 7) short, bluntly hooked at apex, less than half length of subgenital plate. Connective (Figs. 8 & 9) longer than aedeagus.

3 PAUL H. FREYTAG 465 Aedeagus (Figs. 8 & 9) short, without processes, abruptly narrowing to acute apex. Female genitalia: Seventh sternum (Fig. 10) with posterior margin roundly produced on each side, between which it is deeply excavated to median that has two small lobes. Type data Holotype male: Colombia: Amazonas, PNN Amacayacu, San Martin, 3 23 S 70 6 W, 150 m., Malaize, 30-IV-5-V-2000, B. Amado Leg., M694 (IHVL). Paratypes: Two males, same data as holotype, except V-2000, M691 IHVL); one male, same data as holotype, except 5-16-V-2000,M693 (IHVL); one male, same data as holotype, except25-30-v-2000, M695 (IHVL); one male, same data as holotype, except 7-17-VII- 2000, M700 (UKYL); and one male, same data as holotype, except VII-2000, M701 (UKYL). Other specimens seen One male, Colombia: PNN Amacayacu, Mocagua, 3 23 S 70 6 W, 150 m., Malaise, 7-19-VII-2000, A. Parente Leg., M672 (IHVL); two males, same data as holotype, except 24-IX-2-X-2000, M833 (IHVL); one male, same data as holotype, except X-2000, M835 (IHVL); one male, same data as holotype, except 30-VII-8-VIII-2000, M836 (IHVL); one female, same data as holotype, except 24-VIII-1-IX-2000, M837 (IHVL); one male, one female, same data as holotype, except VIII-2000, M842 (IHVL); one male, same data, except Matamata, VIII- 2000, A. Parente Leg., M848 (IHVL); one male, same data as holotype, except XI-2000, M1309 (IHVL); four males, one female, same data as holotype, except 3-11-XI-2000, M1313 (two males, one female-ihvl, two males-ukyl); three males, same data as holotype, except XII- 2000, M1316 (one male-ihvl, two males-ukyl); one female, same data as holotype, except XI-2000, M1318 (IHVL); two females, same data as holotype, except 26-II-12-III-2001, D. Chota, M1611 (IHVL); one male, same data as holotype, except 12-III-2-IV-2001, M1612 (IHVL); one male, same data, except Mocagua, II-2001, M1614 (IHVL); one male, same data, except Matamata, 16-IV-7-V-2001, M1857 (IHVL); one male, same data as holotype, except 3-11-III-2001, M2035 (IHVL); one male, same data as holotype, except 7-11-II-2001, M2038 (IHVL); one female, same data as holotype, except 23-II-3-III-2001, M2040 (IHVL); one male, samd data, except Matamata, 25-VIII-3-IX-2001, D. Chota, M2142 (IHVL); one female, same data, except 3-17-IX-2001, M2241 (IHVL); one male, same data, except 2-15-X- 2001, M2248 (IHVL); one male, same data, except 8-12-III-2000, M. Sharkey (UKYL); one male, one female, Venezuela: T. F. Amazonas, 150 m., Alto Rio Maraca, N W, 14-II-6- III-1989, J. DeMarmela & A. Chacoń (MIZA); and one female, Venezuela: T. F. Amazonas, Dept. Rio Negro, Rio Beria, 140 m., 0 55 N W, 25-III- 15-IV-1984 (MIZA). Notes This species is similar to adspersa in size and color pattern. It differs from adspersa in the shape of the male pygofer process, being triangular shaped in trigona, and sword like in adspersa. The females differ in the shape of the posterior margin of the seventh sternum, being shallowly emarginate in middle with small median lobes intrigona, and deeply but narrowly emarginate in middle in adspersa. Etymology This species is named for the triangular shape of the male pygofer process. Taperinha dozieri Freytag, new species (Figures 11-14) urn:lsid:zoobank.org:act:d251342c a68-b849-f382bccf3b99 Description Length of males 4.5 mm., females unknown. Head with crown yellow, with two large orange spots in posterior half, two small brown slanted spots behind anterior margin near median. Pronotum gray, with large brown patches. Scutellum gray with large brown patches. Forewing opaque gray, with yellow costal margin with six slanting brown veins, saddle of brown near middle. Ventral surface brown. Male genitalia: Pygofer (Fig. 11) longer than wide, pointed at apex, with a ventral process that extends from near base to near apex of pygofer, extending along ventral edge of pygofer, narrowing to pointed apex. Subgenital plate (Figs. 11 & 12) same length as pygofer, triangular in ventral view, bluntly pointed at apex. Style (Fig. 12) triangular at base, with finger-like apex, half length of subgenital plate. Connective (Figs. 13 & 14) same length as aedeagus. Aedeagus (Figs. 13 & 14) nearly same width to apex, slightly curved dorsad,

4 466 TWENTY-ONE NEW SPECIES OF TAPERINHA apex notched. Type data Holotype male: Ecuador: Napa Prov., Yasuni Res. Sta., Rio Tiputini, 250 m., 0 38 S W, at lights, X-1998, Byrd K. Dozier (NMHN). Paratypes: Two males, Ecuador: Provincia de Franciscode Oreliana, Yasuni National Park, S W, 27-IV-2005, ex: sweeping/ night, C. R. Bartlett, N. Nazdrowicz & D. Chang (one-nmnh, one-ukyl). Notes This species is similar to punctata in size and color pattern. The males differ in the placement of the pygofer process, as in dozieri it is parallel to the margin of the pygofer, and in punctata it is not and extends away from the margin of the pygofer.. Etymology This species is named for the collector of the holotype. Taperinha elongata Freytag, new species (Figures 15-18) urn:lsid:zoobank.org:act: f-d4a6-4bec- B39F-BEC21346DBCD Description Length of male 4.1 mm., female 4.5 mm. Head with crown yellow, with two large orange spots on posterior half. Pronotum yellow along anterior margin, with few small orange spots, posterior half dark brown. Scutellum yellow, with median orange spot near apex, triangles brownish orange. Forewings light brown, with distinct brown saddle, costal margin with about six black cross veins. Ventral surface dark brown. Male genitalia: Pygofer (Fig. 15) triangular, bluntly pointed at apex, with a sword-like ventral process, following ventral edge of pygofer, extending just beyond apex of pygofer. Subgenital plate (Figs. 15 & 16) as long as pygofer, triangular in ventral view. Style (Fig. 16) more than half length of subgenital plate, wide at base, abruptly narrowed just beyond middle, than extending as a finger-like apex. Connective (Figs. 17 & 18) shorter than aedeagus. Aedeagus (Figs. 17 & 18) slightly wider at base, then curving dorsad, narrow, long, nearly same width to acute apex. Type data Holotype male: Suriname: Para District, near Overbridge River Resort, 14-II-2010, N W, B. Light, P. Skelly, W. Warner & C. Gillett (FSCA). Paratypes: One female, same data as holotype (FSCA); and one male, Colombia: Vaupés, RN Mosiro-Itajura (Caparú), Igape, 1 4 S W, 60 m., 25-II-4-III-2003, J. Pinzõn, M3627 (IHVL). Notes This species is similar to dozeri in the type of color pattern, but slightly smaller in size. The species differ in that the male pygofer process bends dorsally in dozeri, but bends posteriorly in elongata. Etymology This species is named for the very long and narrow aedeagus. Taperinha stypa Freytag, new species (Figures 19-23) urn:lsid:zoobank.org:act:14c c d7- FD71043F5D2E Description Length of males mm., females mm. Head with crown yellow, two large orange spots in posterior half. Pronotum yellow, with orange band along anterior margin, posterior two thirds yellow brown with orange transverse band on each side of median. Scutellum yellowish brown. Forewing opaque gray, with orange veins, costal margin orange yellow with seven brown slanting veins, brown speckles or spots in most cells, larger brown saddle near middle of wing. Ventral surface, including face, mostly dark brown. Male genitalia: Pygofer (Fig. 19) nearly same width as length, bluntly rounded at apex, with ventral process narrow to apex of pygofer, then wider extending beyond pygofer to near apex of anal tube. Subgenital plate (Figs. 19 & 20) longer than pygofer, triangular in ventral view. Style (Fig. 20) half length of subgenital plate, wide at base, narrowing abruptly to finger-like apex. Connective (Figs. 21 & 22) shorter than aedeagus. Aedeagus (Figs. 21 & 22) spine-like in lateral view, narrowing to acute apex. Female genitalia: Seventh sternum (Fig. 23) twice as wide as long, with posterior margin evenly rounded. Type data Holotype male: Venezuela, Aragua, P.N.H. Pittier, Postachufio, 1100 m., 21- VII-1990, Trampa de Luz, V. Sevini. (MIZA). Paratypes: One female, Venezuela, Aragua, P.N.H. Pittier, 29-X-1992, Rancho Grande, Trampa Interceptacion, J. L. Garcia (MIZA); 1 male, 2 females, Venezeuela, Aragua, Rancho Grande, P. N. Henri Pittier, V-1985, blacklight trap, P.

5 PAUL H. FREYTAG 467 H. Freytag & M. A. Gaiani (UKYL); and one male, same data as last, except 23-VII-1989, at lights, P. H. Freytag & T. Myers (UKYL). Notes This species is near maculata in size and color pattern, with the saddle on the forewings faint and having mostly just a dark black bar near the middle of the wing. Etymology This species is named for the unusual stem to the male pygofer process. Taperinha angulata Freytag, new species (figures 24-28) urn:lsid:zoobank.org:act: b-634e BF8D-D599AE11E31C Description Length of males mm., females mm. Head with crown yellow, with two large reddish orange spots, on each side of median. Pronotum yellow, with orange dash along anterior margin between eyes, also two large orange spots in posterior half. Scutellum yellow, with orange triangles, also orange line down median. Forewings gray, with orange veins, orange brown speckles overall, costal margin orange with five dark brown angled veins. Ventral surface dark brown. Male genitalia: Pygofer (Fig. 24) nearly same width as length, truncate, with sword-like process from ventral margin, curving dorsad then caudad beyond pygofer to near apex of anal tube. Subgenital plate (Figs. 24 & 25) extending half its length beyond pygofer, triangular in ventral view. Style (Fig. 25) two thirds length of subgenital plate, wide at base, narrowing to finger like apex. Connective (Figs. 26 & 27) nearly three times longer than aedeagus. Aedeagus (Figs. 26 & 27) short, bulbous at base, narrowing to acute apex. Female genitalia: Seventh sternum (Fig. 28) twice as wide as long, posterior margin concave on each side of median which extends nearly as long as lateral margins, with shallow emargination in middle. Type data Holotype male: Colombia: Amazonas, PNN Amacayacu, San Martin, 3 23 S 70 6 W, 150 m., Malaise, VIII-2000, B. Amado Leg., M842 (IHVL). Paratypes: one female, same data as holotype, except 2-7-VII-2000, M699 (IHVL); one male same data as holotype, except I-2001, M1311 (IHVL); three males, same as holotype, except 5-13-XII-2000, M1312 (one-ihvl, two-ukyl); 2 males, same data as holotype, except3-11-xi-2000, M1313 (IHVL); one female, same data as holotype, except 29-XII I-2001, M1315 (IHVL); two males, same data as holotype, except, 2-15-I-2001, M1319 (IHVL); one female, same data as holotype, except 30-I-7-II-2001, M2037 (IHVL); and one male, same data as holotype, except 23-II-3-III-2001, M2040 (IHVL). Other specimens seen One male, Colombia: Vaupés, RN Mosiro-Itajura (Capanú), Centro Ambiental, 1 4 S W, 60 m., Malaise, 20-I-1- II-2003, M. Sharkey & D. Arias Leg., M3386 (IHVL); one male, same data, except Antigua Cubaña, 1 4 N 69 3 W, 60 m., 10-III-17-IV-2003, J. Pinzón Leg., M3613 (IHVL); one male, same data, except Centro Ambiental, 1 4 S W, 60 m., 1-9-II-2003, J. Pinzón Leg., M3624 (UKYL); and one male, same data, except Antigua Cubaña, 1 4 N 69 3 W, 60 m., 9-25-II-2003, J. Pinzón Leg., M3629 (IHVL); one male, one female, Ecuador: Provincia de Francisco de Orellana, Yasuni National Park, S W, 220 m., V2 SITE, C. Taylor, HG Vapor Light, 8-I-2010 (UDIC). Notes This species is similar to adspersa in size and the lack of a saddle on the forewings. In angulata the male pygofer process is long, narrow, angling posteriorly beyond the pygofer apex, whereas in adspersa the process is shorter and not extending beyond length of pygofer. Etymology This species is named for the angling male pygofer process. Taperinha bulba Freytag, new species (Figures 29-35) urn:lsid:zoobank.org:act:a5ced2a9-d d-bf9f-15fb085dcb2b Description Length of males mm., females mm. Head with crown yellow, two large orange spots on posterior half, faint line of orange along anterior margin with a pair of dark brown spots near median. Pronotum yellow along anterior margin, with orange band between eyes, posterior two thirds darker yellow brown with a large irregular spot on each side of median. Scutellum yellow, with large orange spots,

6 468 TWENTY-ONE NEW SPECIES OF TAPERINHA Forewings opaque gray, with brown veins, costal margin yellow, with six slanted dark brown veins, otherwise speckled with brown overall, splash of brown near middle. Ventral surface dark brown. Male genitalia: Pygofer (Fig. 29) longer than wide, rounded at apex, with ventral process (Figs. 29 & 30) extending beyond pygofer apex, thin, slightly hooked near acute apex. Subgenital plate (Figs. 29 & 31) longer than pygofer, triangular in ventral view. Style (Fig. 31) half length of subgenital plate, narrowing to blunt finger-like apex. Connective (Figs. 32 & 33) longer than aedeagus. Aedeagus (Figs. 32 & 33) short, bulb-like base, narrowing abruptly to short acute apex, apex (Fig. 34) divided by deep emargination. Female genitalia: Seventh sternum (Fig. 35) twice as wide as long, with posterior margin slightly concave either side of median which is very shallowly emarginate in middle. Type data Holotype male: Peru: Madre de Dios, Rio Tambopata Res., 3 km (air) SW Pto. Maldonado, 290 m., S W, 11-XI-1982, Insect Flight Trap, R. C. Wilkerson (MUSM). Paratypes: Five females, same data as holotype (three-musm, two-ukyl); nine males, four females, Peru: Madre de Dios, Tambopata Res. Zone, Tambopata Res. Cntr. On Rio Tambopata, S W, 192 m., 3-7-X-2004, C. R. Bartlett (five males, 2 females-nmnh, four males, two females-ukyl); and two males, Peru: Madre de Dios, nr Puerto Maldonado, Posadas Amazonas Lodge, at Rio tambopata, 191 m., S W, 30-IX-3-X-2004, C. R. Bartlett (UDIC). Other specimens seen One female, Bolivia: Santa Cruz, 5 km ESE Wernes Hotel, Rio Selva, 20-X-2000, light trap, M. C. Thomas (FSCA); one male, one female, Bolivia: Santa Cruz, Buena Vista, vic. Flora & Fauna Hotel, X-2002, Morris & J. Wappes (UKYL); and two males from Colombia which seem to be this species, one male, Colombia: Tereuta, 26-IV-1967 (OSUC), and one male, Colombia: Choco Dept., Camp Curiche, 1967 (OSUC). Notes This species is also similar to adspersa in size and color pattern. In bulba the male pygofer process angles as in angulata, but is shorter and is hook-shaped in ventral view. The aedeagus is also quite short and bulb-like. Etymology It is named for the bulb-like aedeagus. Taperinha retusa Freytag, new species (Figures 36-42) urn:lsid:zoobank.org:act:48ff9cea-662c B195-D023624F64DB Description Length of males mm., females mm. Head with crown yellow, with pair of small black near median of anterior margin, posterior two thirds with two large orange spots. Pronotum yellow, with orange band across anterior margin between eyes, another orange band across posterior half. Scutellum yellow, with orange speckles. Forewings light brown, with brownish orange speckles overall, costal margin orange, with many black crossveins. Ventral surface very dark brown. Male genitalia: Pygofer (Fig. 36) longer than wide, narrowing to bluntly pointed apex, with ventral process (Figs. 36 & 37) following along ventral margin of pygofer to near apex, slightly curved outward near apex. Subgenital plate (Figs. 36 & 38) longer than pygofer, triangular in ventral view. Style (Fig. 38) half length of subgenital plate, narrowed in middle, with a long acute apex. Connective (Figs. 39 & 40) longer than aedeagus. Aedeagus (Fig. 39 & 40) short, spine-like in lateral view, apex (Fig. 41) divided by short emargination. Female genitalia: Seventh sternum (Fig. 42) twice as wide as long, posterior margin concave on either side of median projection. Type data Holotype male: French Guiana: 33 km SE Roura on Kaw Rd., 18-IV-2007, N W, 227 m., M. V. Light, J. E. Eger (MNHN). Paratypes: Four females, same data as holotype (two-mnhn, two-fsca); one male, one female, same data as holotype, except IV-2007, D. G. Hall & J. E. Eger (FSCA); three females, same data as holotype, except IV-2007, D. G. Hall & J. E. Eger (two- FSCA, one- UKYL). Other specimens seen One female, French Guiana, 17 km W of N2 on Belizon Rd., 3-XII- 2002, 94 m., N W, M. V. Light, J. E. Eger (FSCA); one male, French Guiana, 1 km S Amazone Nature Lodge, 30 km., SE Roura

7 PAUL H. FREYTAG 469 on Kaw Rd., N W, 288M., M. V. Light, 3-4-VI-2005, J. E. ER & M. T. Messenger (UKYL); 1 female, French Guiana, 41 km SE Roura on Kaw Rd., N W, 272 m., 5-7-VI-2005, M. V. Light, J. E. Eger & M. T. Messenger (FSCA); one male, French Guiana, Fourgassie, N W, M. V. Light, J. E. Eger (FSCA); four females, French Guiana, Amazone Nature Lodge, 30 km SE Roura on Kaw Rd., N W, 300 m., , M. V. Light, D. G. Hall & J. E. Eger (FSCA); one male, French Guiana, 33 km SE Roura on Kaw Rd., N, W, 227m., IV- 2007, M. V. Light, D. G. Hall & J. E. Eger (FSCA); and 2 females, French Guiana, Amazone Nature Lodge, N W, 300m., U. V. Light, IV-2007, J. E. Eger (FSCA). Notes This species is also similar to adspersa in size and color pattern. It also has a very short male aedeagus similar to bulba, but not rounded at its base. The male pygofer process is not hooked at its apex, and the style is quite different. Etymology This species is named for the blunt apex of the aedeagus compared to that found in bulba. Taperinha hamata Freytag, new species (Figures 43-46) urn:lsid:zoobank.org:act:6e71ecd9-2d21-4c84-a1a4-f0b70c Description Length of males mm., female unknown. Head with crown yellow, with pair of large orange spots on posterior half. Pronotum yellow across anterior half, with brown spots, posterior half brown. Scutellum yellow, with brown triangles. Forewings light brown, with dark brown saddle, costal margin with many dark brown cross veins. Ventral surface dark brown. Male genitalia: Pygofer (Fig. 43) same length as width, broadly rounded at apex, with two ventral processes; dorsal process following apical margin, than hooking ventrally near apex which extends just beyond apex of pygofer; ventral process sword like extending nearly straight to near apex of dorsal process, apex acute. Subgenital plate (Figs. 43 & 44) longer than pygofer, broad at base, triangular in ventral view. Style (Fig. 44) robust, nearly same length as subgenital plate, tooth like at apex. Connective (Figs. 49 & 50) shorter than aedeagus. Aedeagus slightly curving dorsad near middle, narrowing beyond middle to acute apex, with pair of lateral processes from two thirds length of shaft, paralleling shaft, not extending to apex of aedeagus, but acute at apices. Type data Holotype male: Colombia: Putumayo, PNN La Paya, Cabaña Chagra, 0 7 S W, 320 m., Malaise, XI-2001, R. Cobete Leg., M2572 (IHVL). Paratypes: four males, same data as holotype (two-ihvl, two-ukyl). Other specimens seen One male, same data as holotype, except Mansoya Tierra Baja, 0 6 S W, 330 m., IX-2001, M2073 (IHVL); two males, same data as holotype, except Cabaña La Paya, IX-2001, D. Campos, M2081 (IHVL); two males, same data, except 26-IX-1-X-2001, M2435 (IHVL); three males, same data as holotype, except Fea. Charapa, 0 8 S W, 330 m., X-2001, M2436 (one-ihvl,two-ukyl); one male, same data, except 1-15-X-2001, M2442 (IHVL); one male, same data as holotype, except Cabaña La Paya, 30-XI-15-XII-2001, E. Lozano, M2794 (IHVL); two males, same data except 1-15-XII-2001, E. Lozano, M2795 (IHVL); three males, same data, except XII-2001, M2796 (IHVL); four males, same data, except 20-XI-5-XII-2001, M2798 (two- IHVL, two-ukyl); three males, same data, except Chagra, 30-III-15-IV-2002, R. Cobete Leg., M3149 (IHVL); one male, Colombia: Amazonas, Cgto. La Pedresa, Comunidad Cemaritagua, 1 20 S W, 380 m., Malaise, 24-XII I-2004, J. López Leg., M4168 (IHVL); four males, same data, except M4169 (two-ihvl, two-ukyl); one male, same data, except PNN Amacayacu, Matamata, 3 41 S W, 150 m., A. Perente Leg., M4234 (IHVL); 2 males, same data, except Camino a San Martin, 1-10-III-2004, T. Pape & D. Arias Leg., M4320 (IHVL); one male, Ecuador: Greliana, Reserva Etnica Waorani, 1 km S Onkanegia Camp, S W, 25-VI-1994, fogging terro firme forest, T. L. Erwin, et al (NMNH); one male, Ecuador: Napo Prov., Yasuni Res. Sta., Rio Tipatini, 250 m., 0 38 S W, at light, X-1998, Byrd K. Dozier (FSCA); and one male, Ecuador: Provincia de Francisco de Orellana, Yasuni National

8 470 TWENTY-ONE NEW SPECIES OF TAPERINHA Park, S W, ex Hg Vapor Light/ night, C. R. Bartlett, N. Nazdrowick & D. Chang (NMNH). Notes This species is similar to rugosa in size and color pattern. It differs from rugosa in the apex of the dorsal process of the male pygofer being hooked. In rugosa it is slightly enlarged and triangular and not hooked. Etymology This species is named for the hooked process of the male pygofer. Taperinha media Freytag, new species (Figures 47-50) urn:lsid:zoobank.org:act:385709c8-218f- 4AD4-B0D3-91B688493ADA Description Length of male 4.2 mm., female unknown. Head with crown yellow, with large reddish orange spot on each side of median. Pronotum yellow, with many irregular orange or brown spots. Scutellum yellow, with three wide bands of orange, one down median, one each side, also row of small brown spots across median. Forewings mostly transparent yellow brown, costal area yellow with five angled dark brown veins, median area saddled with darker brown area. Ventral surfaces mostly dark brown. Male genitalia: Pygofer (Fig. 47) long, triangular in lateral view with blunt rounded apex; with two pygofer processes, dorsal process half length of ventral process, angled near base, ventral process just longer than pygofer, slightly sinuate, apex acute. Subgenital plate (Figs ) large in lateral view, triangular in ventral view. Style (Fig. 48) long, nearly straight, with blunt apex. Connective (Figs ) shorter than aedeagus. Aedeagus (Figs ) nearly straight, with pair of lateral short processes just beyond middle of shaft, apex bluntly acute. Type data Holotype male: Colombia: Vaupés, RN Mosiro-Itejura (Caparú), Antigua Cabaña, 1 4 N 69 3 W. 60 m., Malaise, IV- 2003, J. Pinzón Leg., M3615 (IHVL). Notes This species is similar to rugosa in size and color pattern. It has a very unusual aedeagus with lateral processes which is not found in any of the other known species. Etymology This species is named for the medial processes of the aedeagus. Taperinha zanolae Freytag, new species (Figures 51-56) urn:lsid:zoobank.org:act:cd7244c2-f5bc- 43E9-B9D0-0DB44BFA484C Description Length of males mm., females mm. Head with crown light brown, with pair of large dark brown spots on posterior half. Pronotum light brown, with darker brown spots mostly across anterior half. Scutellum brownish orange. Forewings light brown, with brown saddle which is not clearly defined because of other brown speckles in apical half of wing, costal margin orange, with several extra black cross veins. Ventral surface blackish brown. Male genitalia: Pygofer (Figs ) rounded, with two ventral processes, in lateral view processes straight with apices acute, in ventral view slightly curved with ventral process slightly shorter. Subgenital plate (Figs. 51 and 53) large, longer than pygofer in lateral view, triangular in ventral view. Style (Fig. 53) robust at base, with narrowed, laterally curved, blunt apex. Connective (Figs ) short, less than half length of aedeagus. Aedeagus (Figs ) short, gradually narrowing to acute apex, slightly emarginate at apex. Type data Holotype male: French Guiana: 21 km SE Roura on Kaw Rd., 6-7-II-2010, MV Light, N W, J. E. Eger (MNHN). Paratypes: Six males, fifteen females, same data as holotype (two males, five females- MNHN, two males, five females-ukyl, two males, five females-fsca). Other specimens seen Nine males, twenty nine females, French Guiana: Amazonas Nature Lodge, 30 km SE Roura on Kaw Rd., 5-19-II-2010, 300 m., N W, UV light trap, J. E. Eger (five males, twenty five females-fsca, four males, four females-ukyl); six females, same data, except 11-II-2010, MV light, 323 m., N W (FSCA); three females, French Guiana: 33 km SE Roura on Kaw Rd., II-2010, at night, 227 m., N W, J. E. Eger (FSCA); four females, French Guiana: 32 km SE Roura on Kaw Rd., 9-10-II-2010, MV light, 287 m., N W, J. E.

9 PAUL H. FREYTAG 471 Eger (FSCA); one male, one female, same data, except II-2010 (FSCA); seven females, same data, except 13-II-2010 (FSCA); three females, same data, except 18-II-2010 (FSCA); four males, sixteen females, French Guiana: 28 km SE Roura on Kaw Rd., 17-II-2010, MV light, N W, MV light, J. E. Eger (two males, ten females-fsca, two males, six females-ukyl); one male, French Guiana: 21 km SE Roura on Kaw Rd., 16-II-2010, MV light, N W, J. E. Eger (FSCA); two males, one female, Suriname: Para Dist., Zanderij, near guesthouse, N W, at lights, 7-9-II-2010, P. Skelley, W. Wagner & G. Gillett (FSCA); and six males, 2 females, Suriname: Brokopando Dept., Stan Eiland Eco. Resort, near Brownsberg, N W, at lights, II-2010, P. Skelley, W. Wagner & G. Gillett (three males, one female-fsca, three males, one female-ukyl). Notes This species is similar to rugosa in size and color pattern. It differs from that species in the rounded pygofer and the straight pygofer processes. Etymology This species is named for Keti Zanol who has added greatly to our knowledge of this genus. Taperinha undulata Freytag, new species (Figures 57-61) urn:lsid:zoobank.org:act:bf7101a5-958b- 47D FDED97B222A Description Length of males mm., female unknown. Head with crown yellow, two large orange spots on posterior half, also two small brown dashes just posterior to anterior margin near median. Pronotum with yellow and orange across anterior third, posterior two thirds gray with large irregular brownish orange spot covering most of each side. Scutellum yellow, with three large brownish orange spots on anterior half, one orange median spot on posterior half. Forewings transparent light brown, with costal margin yellow with six slanted dark brown veins, median with saddle of darker brown. Ventral surface mostly dark brown. Male genitalia: Pygofer (Fig ) elongate, triangular, with apex bluntly pointed; dorsal process from middle of ventral margin, short, curved dorsad with acute apex, ventral process from base of ventral margin, sinuate, extending to near apex of pygofer. Subgenital plate (Figs. 57 & 59) robust, slightly longer than pygofer, triangular in ventral view. Style (Fig. 59) robust at base, then abruptly narrowing to finger-like apex. Connective (Figs ) equal in length to aedeagus. Aedeagus (Figs ) narrow in ventral view, narrowing to acute apex in lateral view, slight emargination at apex. Type data Holotype male: Colombia: Amazonas, PNN Amacayacu, San Martin, 3 23 S 70 6 W, 150 m., Malaise, XII-2001, D. Chota Leg., M2774 (IHVL). Paratypes: One male, same data as holotype, except Camino a San Martin, 3 41 S W, 1-10-III-2004, T. Pape & D. Arias Leg., M4320 (UKYL); and one male, same data as holotype, except Cabaña Amacayacu, 3 49 S W, 4-10-III-2004, T. Pape & D. Arias Leg., M4323 (IHVL). Notes This species is similar to rugosa in size and color pattern. It differs from that species on several aspects of the male genitalia. The two processes of the male pygofer are quite different in length, and the ventral process is sinuate in ventral view, whereas in rugosa the two processes are equal in length, and not sinuate. Etymology This species is named for the sinuate ventral process of the male pygofer. Taperinha lobata Freytag, new species (Figures 62-65) urn:lsid:zoobank.org:act:eeb bbb- 4F EA673C41FA35 Description Length of males mm., female unknown. Head with crown yellow, two large orange spots on posterior half, two dark brown spots near median of anterior margin, also dark brown around ocelli with extension extending to orange spots on each side. Pronotum yellow, with many spots of brown across anterior third, posterior two thirds brown with many dark brown spots. Scutellum brown, with darker brown spots, posterior half with median orange. Forewings grayish brown, withmany darker brown speckles and spots overall, costal margin yellow with six angled dark brown veins. Ventral surface dark

10 472 TWENTY-ONE NEW SPECIES OF TAPERINHA brown. Male genitalia: Pygofer (Fig. 62) nearly truncate in lateral view, with large concave notch near base on ventral margin; dorsal and ventral processes short, acute at apices, arising from base of posterior margin of pygofer. Subgenital plate (Figs ) long, narrow in lateral view, triangular in ventral view. Style (Fig. 63) large, apical third hook-like. Connective (Figs ) narrow, slightly longer than aedeagus. Aedeagus (Figs ) narrow in ventral view, with bifurcate apex, narrowing to acute apex in lateral view, with bulblike ventral projection at base. Type data Holotype male: Colombia: Putumayo, PNN La Paya, Loma Pendiente, 0 6 S W, 330 m., Malaise, V-2003, L. Megno Leg., M3795 (IHVL). Paratype: One male, same data as holotype, except Cabaña Chagra, 0 7 S W, 320 m., Malaise, XI-2001, R. Cobete Leg., M2572 (UKYL). Notes This species is similar to maculata in size and general color pattern, but in this species the saddle on the forewing is very faint. Also, the male pygofer is emarginate near the base of the ventral margin, and the processes are very short, while in maculata the male pygofer is normal, and the processes very long. The style is large and hook shaped, and in maculata it is smaller and not hook shaped. Etymology This species is named for the unusual shape of the male pygofer. Taperinha stenostyla Freytag, new species (Figures 66-70) urn:lsid:zoobank.org:act:62d3a9d7-1ed0-4e b751f0 Description Length of males mm., female unknown. Head with crown yellow, two large orange spots on posterior half, pair of faint brown angled dashes near median of anterior margin. Pronotum yellow along anterior margin with orange spots, posterior two thirds yellow brown with irregular large brownish orange spot on each side of median. Scutelllum brownish orange on anterior half, posterior half yellow on sides, median orange. Forewings opaque orange brown, with costal margin yellow with six angled dark brown veins, typical dark brown saddle in middle. Male genitalia: Pygofer (Figs ) triangular, with dorsal process arising from near middle of ventral margin, extending posteriorly, acute at apex, ventral process arising from near base of ventral margin, extending to just beyond apex of pygofer, narrow, acute at apex. Subgenital plate (Figs. 66 & 68) robust in lateral view, extending just longer than pygofer, triangular in ventral view. Style (Fig. 68) small, broad at base, with a short finger-like apex. Connective (Figs ) same length as aedeagus. Aedeagus (Figs ) narrow in ventral view, with short bifurcate apex, narrowing to acute apex in lateral view, which curves dorsad. Type data Holotype male: Colombia: Vaupés, RN Mosiro-Itajura (Capanú), Antigua Cabaña, 1 4 N 69 3 W, 60 m., Malaise, 1-9-II-2003, J. Pinzón Leg. M3612 (IHVL). Paratype: one male same data as holotype, except Centro Ambiental, 1 4 S W, 60 m., Red, 20-I-1-II-2003, M. Sharkey & D. Arias Leg., N3387 (UKYL). Other specimen seen One male, Peru: Madre de Dios, Tambopata Res. Zone, Tambopata Res. Centr. On Rio Tambopata, S W, 189 m., 3-7-X-2004, C. R. Bartlett (NMNH). Notes This species is similar to rugosa in size and color pattern. The male pygofer has very narrow, pointed processes, with the dorsal one shorter than the ventral one, and in rugosa the processes are both long, and the ventral one enlarged near the apex. Etymology This species is named for the long narrow ventral process of the male pygofer. Taperinha anisota Freytag, new species (Figures 71-75) urn:lsid:zoobank.org:act:bab81d06-9e de-3bc044078a5f Description Length of males mm., females mm. Head with crown yellow, two large orange spots in posterior half, two dark brown spots near anterior medial margin, also dark brown around ocelli, with extension to orange spots on each side. Pronotum yellow, with many brown spots across anterior third, posterior two thirds brown with many dark brown spots. Scutellum brown, with darker brown spots, posterior median

11 PAUL H. FREYTAG 473 orange. Forewings grayish brown, with many darker brown speckles and spots overall, costal margin yellow, with six angled dark brown veins. Ventral surface dark brown. Male genitalia: Pygofer (Fig. 71) rectangular in lateral view, with large notch near base of ventral margin; both processes from base of posterior margin, dorsal process very short, acute at apex, ventral process narrow, extending nearly to apex of pygofer, acute at apex. Subgenital plate (Figs ) narrow in lateral view, longer than pygofer, triangular in ventral view. Style (Fig.72) large, with large finger-like hook at apex. Connective (Figs ) narrow, same length as aedeagus. Aedeagus (Figs ) narrowing slightly in ventral view to slightly incised apex, in lateral view very narrow beyond base to acute apex, with bump-like process on ventral side of base. Female genitalia: Seventh sternum with posterior margin excavated on each side to median half way to base, with finger-like projection in middle of excavation extending posterior to length of sternal segment. Type data Holotype male: Colombia: Amazonas, PNN Amacayacu, San Martin, 3 23 S 70 6 W, 150 m., Malaise, 24-VIII-1-IX-2000, B. Amado Leg., M837 (IHVL). Paratypes: One female, same data as holotype, except 24-IX-2-X-2000, M833 (IHVL); one male, same data as holotype, except 2-10-X-2000, M843 (UKYL); one male, one female, same data as holotype, except XI- 2000, M1309 (UKYL); and one male, same data as holotype, except 29-XII I-2001, M1315 (IHVL). Notes This species is similar to lobata in size and color pattern. It differs from that species in the very short curved dorsal process of the male pygofer. It also differs in the aedeagus having a very shallow emargination at the apex of the aedeagus, whereas in lobata the apex has a much deeper emargination. Etymology This species is named for the unusual shape of the aedeagus in lateral view. Taperinha lapayensis Freytag, new species (Figures 76-79) urn:lsid:zoobank.org:act:2f62082b-4bcd B2F1-91F2022C6B3A Description Length of males 4.5 mm., female 4.9 mm. Head with crown yellow, two large brownish orange spots on posterior half, two small slanting dark brown dashes near anterior margin medially, dark brown around ocelli with extension to large brownish orange spot on each side. Pronotum with anterior third yellow, with large brown spots, posterior two thirds grayish brown with large irregular brown spot on each side of median. Scutellum with anterior half brown with large brownish orange spots, posterior half with sides yellow, median orange. Forewings mostly opaque grayish brown, median with large brown spot, between this spot and posterior two thirds of clavus darker brown spots, costal margin yellow with six slanting dark brown veins. Ventral surface dark brown. Male genitalia: Pygofer (Fig. 76) robust, nearly rounded at apex, with two short processes arising from base of posterior margin, dorsal process extending to near dorsal margin of pygofer, ventral process curving dorsad away from pygofer, both acute at apices. Subgenital plate (Figs ) robust, extending beyond pygofer in lateral view, triangular in ventral view. Style (Fig. 77) with broad base, narrowing to bent finger-like apex. Connective (Figs ) narrow, same length as aedeagus. Aedeagus (Figs ) short, narrowing to slightly notched apex, curving dorsad in lateral view, narrowing to acute apex, with expanded bump-like base. Type data Holotype male: Colombia: Putumayo, PNN La Paya, Cabaña La Paya, Chagra, 0 7 S W, 320 m., Malaise, III-2002, R. Cobete Leg., M3148 (IHVL). Paratypes: One male, one female, same data as holotype (male-ukyl, female-ihvl). Other specimens seen One male, Ecuador: Provincia de Francisco de Orellana, Yasuni National Park, S W, 220 m., V2 SITE, C. Taylor, HG Vapor Light, 8-I-2010 (UDIC); and one female, same data, except 7-I-2010 (UDIC). Notes This species is similar to rugosa in

12 474 TWENTY-ONE NEW SPECIES OF TAPERINHA size and color pattern. It differs from that species by having the processes of the male pygofer coming off the pygofer together. Etymology This species is named for the locality of the holotype. Taperinha odonta Freytag, new species (Figures 80-84) urn:lsid:zoobank.org:act:216eebf7-012e-43d3- B044-08C8C777FFEC Description Length of males mm., female unknown. Head with crown yellow, with two large orange spots in posterior half. Pronotum with anterior narrowly yellow with brownish orange spots, posterior mostly grayish brown with brown spots. Scutellum yellow, with anterior half with three large brownish orange spots, posterior with large median orange spot. Forewings mostly opaque brownish orange, with costal margin yellow with six slanted dark brown veins, median saddle of darker brown. Ventral surface dark brown. Male genitalia: Pygofer (Figs ) triangular in lateral view, with two processes from base of ventral margin, dorsal process on median side of pygofer bending toward middle of genital capsule, acute at apex, ventral process narrow, extending nearly length of pygofer, apex acute. Subgenital plate (Figs. 80 & 82) long, narrow in lateral view, triangular in ventral view. Style (Fig. 82) robust base, narrowing abruptly to finger-like apex, which appears tooth-like at apex. Connective (Figs ) narrow, shorter than aedeagus. Aedeagus (Figs ) narrow in ventral view, narrowing to blunt apex in lateral view. Type data Holotype male: Colombia: Vaupés, RN Mosiro-Itajura (Caparú), Centro Ambiental, 1 4 S W, 60 m., Malaise, 20-I-1- II-2003, M. Sharkey & D. Arias Leg., M3386 (IHVL). Paratypes: Two males, same data as holotype (one-ihvl, one UKYL); and one male, same data as holotype, except Red, M3387 (IHVL). Other specimens seen One male, same data as holotype, except Igapo, 7-22-X-2002, L. Benavides Leg., M3395 (IHVL); one male, same data except 2-22-XI-2002, M3397 (IHVL); one male, same data as holotype, except Antigua Cabaña, 1 4 S 69 3 W, III-2003, J. Pinzón, M3611 (IHVL); one male same data as holotype, except 27-III-3-IV-2003, J. Pinzón, M3620 (IHVL); two males, same data as holotype, except III- 2003, L. Benavides, M3626 (UKYL); and one male, Colombia: Amazonas, PNN Amacayacu, Metameta, 3 41 S W, 150 m., Red, 26-II-2001, D. Chota Leg., M1632 (IHVL). Notes This species is similar to rugosa in size and color pattern. It has quite different processes on the male pygofer. In odonta they are both diverging and narrowing to acute apices, and in rugosa they are close together, and have different apices. Etymology This species is named for the tooth like apex to the style. Taperinha peruviana Freytag, new species (Figures 85-89) urn:lsid:zoobank.org:act:782b3c56-a2f b57c-adfbbbcbe56b Description Length of male 4.6 mm., female unknown. Head with crown yellow, two large orange spots in posterior half, faint small brown slanting dashes near anterior margin medially. Pronotum yellow along anterior margin with orange spots, posterior two thirds darker gray brown with large irregular brownish orange spots. Scutellum yellow, with large orange spots. Forewings opaque gray, with seven slanting dark brown veins along costal margin, with dark brown saddle in middle. Ventral surface brown. Male genitalia: Pygofer (Figs ) triangular in lateral view, with two long processes arising from thickened ventral margin, dorsal process on median side of pygofer curving posteriorly to end near apex of pygofer, apex acute, ventral process extending along posterior margin of pygofer to just beyond apex of pygofer, thicker in ventral view with acute apex. Subgenital plate (Figs. 85 & 87) narrow, extending to length of pygofer, triangular in ventral view. Style (Fig. 87) small, robust at base, apex short finger-like, bluntly pointed. Connective (Figs ) narrow, same length as aedeagus. Aedeagus (Figs ) narrow, slightly expanded near base in ventral view, divided at apex, narrowing to sharply acute apex in lateral view, base with bulge at base. Type data Holotype male: Peru: Madre de Dios, Tambopata Res. Zone, Tambopata Res. Centr.

13 PAUL H. FREYTAG 475 On rio Tambopata, S W, 189 m.,3-7-x-2004, C. R. Bartlett (MUSM). Notes This species is similar to odonta in size and color pattern. It differs by having the dorsal process of the male pygofer curving toward the ventral process in lateral view. Etymology This species is named for the country from which it was collected. Taperinha longicornis Freytag, new species (Figures 90-93) urn:lsid:zoobank.org:act:761c70cc c1d-b16c-8b29c23817b9 Description Length of male 5.0 mm., female unknown. Head with crown yellow, with two large orange spots in posterior half. Pronotum grayish yellow, with two brownish orange large spots. Scutellum mostly brownish orange. Forewing opaque gray, with costal margin yellow with five dark brown slanting veins, median with large dark brown spot, with faint indication of saddle of dark brown spots. Ventral surface brown. Male genitalia: Pygofer (Fig. 90) rectangular in lateral view, with two short processes from posterior margin, dorsal process from middle of margin curving slightly posterior, with acute apex, ventral process from base of posterior margin curving dorsad, with acute apex. Subgenital plate (Figs ) extending beyond pygofer, with apex bent dorsad, triangular in ventral view. Style (Fig. 91) base stout, long, with narrow, short finger-like apex. Connective (Figs ) narrow, half length of aedeagus. Aedeagus (Figs ) narrow, very long, divided into two pointed processes at apex, curving dorsad in lateral view. Type data Holotype male: Ecuador: Napa Prov., Yasuni Res. Sta., Rio Tiputini, 250 m., 7 38 S W, at lights, X-1998, Byrd K. Dozier (NMNH). Notes This species is similar to lapayensis in size and general color pattern. It differs by having a very long stem to the aedeagus, with a deep emargination at the apex. Etymology This species is named for the very long aedeagus. Taperinha Linnavuori Taperinha Linnavuori 1959, species Type species Taperinha bifurcata Linnavuori adspersa Linnavuori & DeLong 1978, 131 Bolivia, Ecuador, Peru albinoi Zanol 1988, 415 Brazil, Peru angulata Freytag n. sp. Colombia anisota Freytag n. sp. Colombia assurinia Zanol 2004, 340 Brazil bifurcata Linnavuori 1959, 190. Brazil, Bolivia, Peru brunneopicta Zanol 1988, 412. Brazil, Ecuador, Peru, French Guiana bulba Freytag n. sp. Peru, Colombia compta Zanol 2004, 338. Brazil concinna Zanol 2004, 338. Brazil, Colombia, Venezuela dentata Zanol 1988, 415 Brazil discigutta Linnavuori 1959, 189 Brazil dozieri Freytag n. xp. Ecuador elongata Freytag n. sp. Colombia, Peru excavata Zanol 1989, 256. Guyana, Suriname, Colombia flavescens Zanol 1988, 412. Brazil freytagi Zanol 2004, 341. Brazil hamata Freytag n. sp. Colombia kijaua Zanol 2004, p Brazil lapayensis Freytag n. sp. Colombia lobata Freytag n. sp. Colombia longicornis Freytag n. sp. Ecuador maculata Zanol 1989, 255. Peru manauara Zanol 2004, 337. Brazil marinonii Zanol 1997, 625. Brazil media Freytag n. sp. Colombia monticola Linnavuori & DeLong 1978, 131 Peru nuda Freytag n. sp. Colombia odonta Freytag n. sp. Colombia ourensis Zanol 2004, 342 Brazil peruviana Freytag n. sp. Peru punctata Linnavuori 1959, 190 Costa Rica, Panama, Nicaragua, Colombia, Ecuador pupejua Zanol 2004, 341 Brazil, Peru retusa Freytag n. sp. French Guiana rugosa Zanol 1988, 411. Brazil